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hairy lip fern

Cleveland's lip fern

Stems

compact to short-creeping, usually 4–8 mm diam.;

scales often uniformly brown but at least some on each plant with thin, poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent.

usually short-creeping, 1–3 mm diam.;

scales usually bicolored, with well-defined, dark, central stripe and broad, light brown margins, lanceolate, straight to slightly contorted, strongly appressed, persistent.

Leaves

clustered, 7–50 cm;

vernation circinate.

clustered to somewhat scattered, 8–40 cm;

vernation noncircinate.

Petiole

dark brown, rounded adaxially.

dark to light brown, rounded adaxially.

Blade

linear-oblong to lanceolate, usually 2-pinnate-pinnatifid at base, 1.5–5 cm wide;

rachis rounded adaxially, lacking scales, with monomorphic pubescence.

oblong-lanceolate to ovate, usually 4-pinnate at base, 2–8 cm wide;

rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence.

Ultimate segments

oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially and adaxially sparsely hirsute with long, segmented hairs.

round to subcordate, beadlike, the largest 1–2 mm, abaxially with branched hairs and small ciliate scales, adaxially glabrous.

Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing sparsely hirsute adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

marginal, weakly differentiated, 0.05–0.25 mm wide.

Sori

discontinuous, concentrated on small apical and lateral lobes.

± continuous around segment margins.

Sporangia

containing 64 spores.

containing 64 spores.

Costae

brown adaxially for most of length;

abaxial scales absent.

green adaxially for most of length;

abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, 0.4–1 mm wide, imbricate, occasionally concealing ultimate segments, ciliate (usually only on proximal 1/2).

2n

= 60.

Cheilanthes lanosa

Cheilanthes clevelandii

Phenology Sporulating summer–fall. Sporulating late spring–summer.
Habitat Rocky slopes and ledges, on a variety of substrates including limestone and granite Rocky slopes and ledges, usually on igneous substrates
Elevation 100–800 m (300–2600 ft) 0–1600 m (0–5200 ft)
Distribution
from FNA
AL; AR; CT; FL; GA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV
from FNA
CA; Mexico in Baja; California
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Discussion

Cheilanthes lanosa is apparently confined to the forests and prairies of eastern North America, and reports of this distinctive species from Arizona and New Mexico (A. J. Petrik-Ott 1979) have not been substantiated by herbarium specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although some specimens of Cheilanthes clevelandii can be difficult to distinguish from C. covillei and C. intertexta (see comments under C. covillei), the species is restricted to the coastal mountains of California and Baja California, and it rarely overlaps the ranges of these closely related species. In the region where C. clevelandii is sympatric with C. intertexta, the smaller spores of the former species (averaging less that 55 µm in diameter) are helpful in identification. T. Reeves (1979) tentatively identified two varieties of C. clevelandii, but formal recognition of these taxa must await further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms Nephrodium lanosum, C. vestita
Name authority (Michaux) D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2(1): 234. (1859) D. C. Eaton: Bull. Torrey Bot. Club 6: 33. (1875)
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