Cheilanthes intertexta |
Cheilanthes viscida |
|
---|---|---|
coastal lip fern |
viscid lip fern |
|
Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping, usually 4–8 mm diam.; scales uniformly brown, linear-subulate, strongly contorted, loosely appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 6–30 cm; vernation circinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, flattened or slightly grooved distally on adaxial surface. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
narrowly oblong to linear, 3-pinnate-pinnatifid at base, 1–4 cm wide; rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
oblong to lanceolate, not beadlike, the largest 3–4 mm, abaxially and adaxially glandular-pubescent with short, sticky, capitate glands. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing glandular pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
usually discontinuous, concentrated on apical and lateral lobes. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
green adaxially for most of length; abaxial scales absent. |
Cheilanthes intertexta |
Cheilanthes viscida |
|
Phenology | Sporulating late spring–fall. | Sporulating late spring–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Cliffs and rocky slopes, usually on igneous substrates |
Elevation | 500–2800 m (1600–9200 ft) | 200–1300 m (700–4300 ft) |
Distribution |
CA; NV; OR
|
CA; Mexico in Baja California
|
Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes viscida is confined to a relatively small region in the deserts of California. Variations in spore size among populations suggest that the species may include more than one cytotype. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | Davenport: Bull. Torrey Bot. Club 6: 191. (1877) |
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