Cheilanthes intertexta
Cheilanthes tomentosa
coastal lip fern
woolly lip fern
short-creeping, usually 3–7 mm diam.;
scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent.
compact, usually 4–8 mm diam.;
scales mostly bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent.
clustered, 4–25 cm;
vernation noncircinate.
clustered, 8–45 cm;
vernation noncircinate.
dark brown, rounded adaxially.
usually dark brown, rounded adaxially.
lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide;
rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence.
oblong-lanceolate, usually 4-pinnate at base, 1.5–8 cm wide;
rachis rounded abaxially, with scattered linear scales and monomorphic pubescence.
oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent.
oval or rarely oblong, beadlike, the largest 1–2 mm, abaxially densely tomentose, adaxially pubescent with fine, unbranched hairs.
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially.
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing tomentose adaxially.
marginal, weakly differentiated, 0.05–0.25 mm wide.
marginal to obscurely inframarginal, somewhat differentiated, 0.05–0.25 mm wide.
± continuous around segment margins.
± continuous around segment margins.
containing 64 spores.
containing 32 spores.
green adaxially for most of length;
abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2.
green adaxially for most of length;
abaxial scales multiseriate, linear, truncate at base, inconspicuous, the largest 0.1–0.4 mm wide, loosely imbricate, not concealing ultimate segments, usually entire, not ciliate.
= 2n = 90, apogamous.
Cheilanthes intertexta
Cheilanthes tomentosa
Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Cheilanthes tomentosa is an apogamous triploid of unknown parentage. It is closely related to C. eatonii, but it is distinguished by having narrower, less prominent costal scales. Natural hybrids between these two species have been reported (D. S. Correll 1956), but such hybrids are unlikely because both species are apogamous in the supposed region of hybridization.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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