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coastal lip fern

Pringle's lip fern

Stems

short-creeping, usually 3–7 mm diam.;

scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent.

long-creeping, 1–3 mm diam.;

scales uniformly brown or with poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, usually persistent.

Leaves

clustered, 4–25 cm;

vernation noncircinate.

clustered to somewhat scattered, 4–15 cm;

vernation noncircinate.

Petiole

dark brown, rounded adaxially.

dark brown, grooved distally on adaxial surface.

Blade

lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide;

rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence.

ovate-deltate, 3-pinnate-pinnatifid at base, 1.5–5 cm wide;

rachis grooved adaxially, with scattered, lanceolate scales, not pubescent.

Ultimate segments

oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent.

spatulate, not especially beadlike, the largest usually 2–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous.

Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair conspicuously larger than adjacent pair, inequilateral, basiscopic pinnules enlarged, appearing glabrous adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

marginal, weakly differentiated, 0.05–0.25 mm wide.

Sori

± continuous around segment margins.

discontinuous, confined to apical or lateral lobes.

Sporangia

containing 64 spores.

containing 64 spores.

Costae

green adaxially for most of length;

abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2.

green adaxially for most of length;

abaxial scales multiseriate, lanceolate, truncate or subcordate at base, without overlapping basal lobes, conspicuous, the largest 0.4–0.8 mm wide, loosely imbricate, not concealing ultimate segments, erose, not ciliate.

2n

= 60.

Cheilanthes intertexta

Cheilanthes pringlei

Phenology Sporulating late spring–fall. Sporulating late spring–fall.
Habitat Rocky slopes and ledges, usually on igneous substrates Rocky slopes and ledges, usually on igneous substrates
Elevation 500–2800 m (1600–9200 ft) 700–1200 m (2300–3900 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
from FNA
AZ; n Mexico
Discussion

Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cheilanthes pringlei is often confused with young, sterile plants of C. fendleri, but it is easily distinguished from the latter by having rachises that are grooved adaxially. This species appears to be restricted to the Sonoran Desert; records from Gila and Cochise counties, Arizona, and southern New Mexico are based on misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms C. covillei subsp. intertexta
Name authority (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) Davenport: Bull. Torrey Bot. Club 10: 61. (1883)
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