Cheilanthes intertexta |
Cheilanthes newberryi |
|
---|---|---|
coastal lip fern |
Newberry's lip fern |
|
Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping to compact, usually 3–5 mm diam.; scales uniformly dark brown to black or with light brown, ephemeral tip, linear-subulate, slightly contorted, strongly appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 5–30 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
lanceolate to linear-oblong, 2-pinnate-pinnatifid to 3-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, lacking scales, with dense monomorphic pubescence. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially densely woolly with very fine, cobwebby hairs, tomentose adaxially. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, proximal basiscopic pinnules slightly enlarged, appearing tomentose adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
absent. |
Sori | ± continuous around segment margins. |
usually discontinuous, concentrated on small apical and lateral lobes. |
Sporangia | containing 64 spores. |
containing 64 or 32 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
green or straw-colored adaxially for most of length; abaxial scales absent. |
2n | = 60. |
|
Cheilanthes intertexta |
Cheilanthes newberryi |
|
Phenology | Sporulating late spring–fall. | Sporulating late spring–summer. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Rocky cliffs and slopes, usually on igneous substrates |
Elevation | 500–2800 m (1600–9200 ft) | 0–1200 m (0–3900 ft) |
Distribution |
CA; NV; OR
|
CA; Mexico in Baja California
|
Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As with Cheilanthes bonariensis, C. newberryi has traditionally been assigned to Notholaena; R. M. Tryon and A. F. Tryon (1982) argued for its placement in Cheilanthes. Hybridization of C. newberryi with C. covillei to form C. × fibrillosa supports placement in Cheilanthes. R. M. Tryon (1956) reported variations in spore number per sporangium in C. newberryi that may indicate cytologic variability within the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | Notholaena newberryi |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | (D. C. Eaton) Domin: Biblioth. Bot. 20(85): 133. (1915) |
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