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coastal lip fern

fairy swords

Stems

short-creeping, usually 3–7 mm diam.;

scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent.

long-creeping, 0.7–3 mm diam.;

scales uniformly brown, ovate-lanceolate, straight to slightly contorted, loosely appressed, often deciduous on older portions of stem.

Leaves

clustered, 4–25 cm;

vernation noncircinate.

scattered, 7–30 cm;

vernation noncircinate.

Petiole

dark brown, rounded adaxially.

usually dark brown, rounded adaxially.

Blade

lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide;

rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence.

oblong-lanceolate to ovate-deltate, 4-pinnate at base, 2–5 cm wide;

rachis rounded adaxially, with scattered linear-lanceolate scales and sparse monomorphic pubescence.

Ultimate segments

oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent.

round to slightly oblong, beadlike, the largest 0.7–1 mm, abaxially nearly glabrous, often with a few small scales or branched hairs, adaxially appearing tomentose but actually nearly glabrous.

Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing densely tomentose adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

marginal, weakly differentiated, 0.05–0.25 mm wide.

Sori

± continuous around segment margins.

± continuous around segment margins.

Sporangia

containing 64 spores.

containing 32 spores.

Costae

green adaxially for most of length;

abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2.

green adaxially for most of length;

abaxial scales multiseriate, lanceolate-ovate, truncate to cordate at base, usually without overlapping basal lobes, conspicuous, the largest 0.4–1 mm wide, strongly imbricate, often concealing ultimate segments, long-ciliate throughout, cilia fine, curly, forming entangled mass.

n

= 2n = 90, apogamous.

Cheilanthes intertexta

Cheilanthes lindheimeri

Phenology Sporulating late spring–fall. Sporulating summer–fall.
Habitat Rocky slopes and ledges, usually on igneous substrates Rocky slopes and ledges, on a variety of acidic to mildly basic substrates
Elevation 500–2800 m (1600–9200 ft) 200–2500 m (700–8200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
from FNA
AZ; NM; TX; Mexico
Discussion

Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cheilanthes lindheimeri is an apogamous triploid of unknown parentage. It is occasionally misidentified as C. wootonii (actually the element here recognized as C. yavapensis); most specimens can be placed using the characteristics given in the key. The adaxial blade surface appears to be densely gray tomentose, but this is an illusion created by the fine, curly cilia of the abaxial costal scales that overtop the minute ultimate segments and form an entangled mass that prevents the easy removal of individual costal scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms C. covillei subsp. intertexta Myriopteris lindheimeri
Name authority (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) Hooker: Sp. Fil. 2: 101, plate 107a. (1852)
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