Cheilanthes intertexta |
Cheilanthes horridula |
|
---|---|---|
coastal lip fern |
prickly lip fern, rough lipfern |
|
Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping, usually 4–7 mm diam.; scales uniformly brown or with poorly defined, dark central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 5–30 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
black to dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
linear-oblong to lanceolate, pinnate-pinnatifid to 2-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered linear-lanceolate scales and dimorphic pubescence, abaxially sparsely hirsute, adaxially covered with tortuous, appressed hairs. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
narrowly elliptic to elongate-deltate, not beadlike, the largest 3–5 mm, abaxially and adaxially scabrous with stiff, usually pustulose hairs. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing pustulose adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, slightly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
green adaxially for most of length; abaxial scales multiseriate, lanceolate, truncate to subcordate at base, without overlapping basal lobes, somewhat inconspicuous, the largest 0.4–0.6 mm wide, loosely imbricate, not concealing ultimate segments, erose, not ciliate. |
2n | = 58, 116. |
|
Cheilanthes intertexta |
Cheilanthes horridula |
|
Phenology | Sporulating late spring–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Rocky slopes and ledges, usually on limestone |
Elevation | 500–2800 m (1600–9200 ft) | 100–1400 m (300–4600 ft) |
Distribution |
CA; NV; OR
|
OK; TX; n Mexico |
Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The scabrous, pustulose hairs of Cheilanthes horridula make it one of the most distinctive species of Cheilanthes in North America. As currently circumscribed, the species includes two sexually reproducing cytotypes that may be given formal recognition when their morphologic characteristics and distributions are sufficiently well known. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | Pellaea aspera |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | Maxon: Amer. Fern J. 8: 94. (1918) |
Web links |