Cheilanthes intertexta |
Cheilanthes gracillima |
|
---|---|---|
coastal lip fern |
lace fern, lace lipfern |
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Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping, 4–8 mm diam.; scales uniformly brown or with poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 5–25 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
linear-oblong, 2–3-pinnate at base, 1–2.5 cm wide; rachis rounded adaxially, with scattered linear scales, not pubescent. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
oblong or rarely oval, beadlike, the largest 1.5–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered, branched hairs or glabrescent. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing sparsely pubescent or glabrous adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, slightly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
green adaxially for most of length; abaxial scales multiseriate, linear, truncate at base, inconspicuous, the largest 0.1–0.4 mm wide, loosely imbricate, not concealing ultimate segments, long-ciliate, cilia usually confined to base. |
Cheilanthes intertexta |
Cheilanthes gracillima |
|
Phenology | Sporulating late spring–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Cliffs and rocky slopes, usually on igneous substrates |
Elevation | 500–2800 m (1600–9200 ft) | 800–3000 m (2600–9800 ft) |
Distribution |
CA; NV; OR
|
CA; ID; MT; NV; OR; UT; WA; AB; BC
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Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes gracillima is a well-marked species, but it apparently hybridizes with C. intertexta (see reticulogram) to produce plants of intermediate morphology with malformed spores that have been called C. gracillima var. aberrans M. E. Jones (A. R. Smith 1974). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | Myriopteris gracillima |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2: 234. (1859) |
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