Cheilanthes intertexta |
Cheilanthes covillei |
|
---|---|---|
coastal lip fern |
Coville's lip fern |
|
Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping, usually 2–4 mm diam.; scales usually uniformly dark brown to black or rarely with narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 5–30 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
lanceolate to ovate-deltate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, somewhat scaly, not pubescent. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
round to oblong, beadlike, the largest 1–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous (or somewhat scaly) adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, usually concealing ultimate segments, ciliate only on basal lobes. |
2n | = 60. |
|
Cheilanthes intertexta |
Cheilanthes covillei |
|
Phenology | Sporulating late spring–fall. | Sporulating late spring–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Rocky slopes, cliffs, and ledges, usually on igneous substrates |
Elevation | 500–2800 m (1600–9200 ft) | 100–2500 m (300–8200 ft) |
Distribution |
CA; NV; OR
|
AZ; CA; NV; UT; Mexico in Baja California
|
Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes covillei can be difficult to distinguish from the closely related C. intertexta and C. clevelandii; it differs from these two species in having glabrous blades and costal scales ciliate only on the basal lobes. Cheilanthes covillei is occasionally misidentified as C. fendleri because the cilia of the scales are often obscure; it is distinguished from the latter species by having rigid, dark brown stem scales that are strongly appressed. Cheilanthes covillei hybridizes with C. parryi and C. newberryi to form rare, sterile diploids known as C. × parishii Davenport and C. × fibrillosa (Davenport) Davenport ex Underwood, respectively. A third sterile diploid hybrid with C. fendleri has recently been discovered in central Arizona (M. D. Windham, unpublished). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | Myriopteris covillei |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | Maxon: Proc. Biol. Soc. Wash. 31: 147. (1918) |
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