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coastal lip fern

Arizona lip fern

Stems

short-creeping, usually 3–7 mm diam.;

scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent.

short-creeping, usually 4–8 mm diam.;

scales uniformly dark brown or with poorly defined, dark, central stripe, narrowly lanceolate, straight to slightly contorted, loosely appressed, usually persistent.

Leaves

clustered, 4–25 cm;

vernation noncircinate.

clustered, 8–25 cm;

vernation circinate.

Petiole

dark brown, rounded adaxially.

dark brown, grooved most of length adaxially.

Blade

lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide;

rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence.

elongate-pentagonal, 3-pinnate-pinnatifid to 4-pinnate at base, 4–10 cm wide;

rachis grooved adaxially, lacking scales, with minute, widely scattered hairs.

Ultimate segments

oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent.

elliptic to linear-oblong, not beadlike, the largest 4–10 mm, abaxially with reddish, scattered, sessile glands, adaxially glabrous;

false indusia marginal to obscurely inframarginal, strongly differentiated, 0.25–0.50 mm wide.

Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair conspicuously larger than adjacent pair, inequilateral, proximal basiscopic pinnules enlarged, glabrous adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

Sori

± continuous around segment margins.

± continuous around segment margins.

Sporangia

containing 64 spores.

containing 32 spores.

Costae

green adaxially for most of length;

abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2.

green adaxially for most of length;

abaxial scales absent.

n

= 2n = 90, apogamous.

Cheilanthes intertexta

Cheilanthes arizonica

Phenology Sporulating late spring–fall. Sporulating summer–fall.
Habitat Rocky slopes and ledges, usually on igneous substrates Rocky slopes and ledges, on quartzite or igneous substrates
Elevation 500–2800 m (1600–9200 ft) 1900–2400 m (6200–7900 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
from FNA
AZ; Mexico; Central America in Guatemala; Honduras
Discussion

Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The distinctive Cheilanthes arizonica is an apogamous triploid of unknown parentage. In the flora, it is known from about seven localities, all of which are situated within 50 km of the Mexican border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms C. covillei subsp. intertexta C. pyramidalis subsp. arizonica, C. pyramidalis var. arizonica
Name authority (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) (Maxon) Mickel: Phytologia 41: 433. (1979)
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