Cheilanthes intertexta |
Cheilanthes aemula |
|
---|---|---|
coastal lip fern |
rival lipfern, Texas lip fern |
|
Stems | short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
short-creeping, usually 4–7 mm diam.; scales uniformly brown or slightly darker at base, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
Leaves | clustered, 4–25 cm; vernation noncircinate. |
clustered, 10–50 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
black to dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
ovate-deltate, 3-pinnate to 3-pinnate-pinnatifid at base, 4–15 cm wide; rachis rounded adaxially, lacking scales, with dimorphic pubescence, abaxially sparsely hirsute, adaxially covered with tortuous, appressed hairs. |
Ultimate segments | oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
narrowly elliptic to elongate-deltate, not beadlike, the largest 3–6 mm, abaxially and adaxially sparsely hirsute to glabrescent. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair slightly larger than adjacent pair, somewhat inequilateral, proximal basiscopic pinnules conspicuously enlarged, appearing glabrous or sparsely pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, slightly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
somewhat discontinuous, often concentrated on interrupted lateral lobes. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
black adaxially for most of length; abaxial scales absent. |
2n | = 58. |
|
Cheilanthes intertexta |
Cheilanthes aemula |
|
Phenology | Sporulating late spring–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Rocky slopes and ledges, apparently confined to limestone |
Elevation | 500–2800 m (1600–9200 ft) | 100–500 m (300–1600 ft) |
Distribution |
CA; NV; OR
|
TX; n Mexico |
Discussion | Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the characteristics mentioned in the key, Cheilanthes aemula is distinguished from North American populations of C. alabamensis by having 64 spores per sporangium rather than 32. In the flora, this species is known from about 10 localities in central and western Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | |
Name authority | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) | Maxon: Contr. U.S. Natl. Herb. 10: 495. (1908) |
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