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slender chasmanthium, slender woodoats, spike uniola

Habit Plants annual or perennial; rhizomatous or stoloniferous.
Culms

40-130 cm, to 1 mm thick at the nodes, unbranched, leafy for 50% of their height.

annual, sometimes becoming woody, internodes solid or hollow.

Sheaths

glabrous;

collars glabrous;

ligules 0.2-0.4 mm, entire;

blades (8)15-35(40) cm long, 3-8(11) mm wide, linear-lanceolate, usually glabrous, sometimes sparsely pilose adaxially.

Leaves

distichous;

sheaths usually open;

auricles sometimes present;

abaxial ligules absent or of hairs;

adaxial ligules membranous, ciliate or not, or of hairs;

blades often pseudopetiolate;

mesophyll cells non-radiate;

adaxial palisade layer often present;

fusoid cells absent, but fusoidlike cells frequently present as extensions of the outer parenchyma bundle sheath;

arm cells absent;

kranz anatomy absent;

midrib simple;

adaxial bulliform cells present, large;

stomata with dome-shaped or triangular subsidiary cells;

bicellular microhairs present, with long, tapering apical cells;

papillae absent.

Panicles

(7)12-35(47) cm, contracted, erect;

branches ascending to appressed;

axils of panicle branches glabrous;

pedicels 0.5-2.5 mm.

Inflorescences

ebracteate, racemose or paniculate, panicle branches sometimes spikelike;

disarticulation below the spikelets or the florets, sometimes at the base of the pedicels.

Spikelets

4-9 mm long, 2-6 mm wide, with (2)3-5(7) florets, lower 1(2) florets sterile, fertile florets divergent to 45°.

bisexual or unisexual, often laterally compressed, with (1)2-many florets, reduced florets present, distal or basal to the functional florets.

Glumes

shorter than the lemmas;

lemmas lacking uncinate hairs, usually 5-9-veined, unawned or with single, terminal awns;

paleas usually well-developed, sometimes short compared to the lemmas;

lodicules 2 or none, cuneate, usually well-vascularized, varying to not or scarcely vascularized;

stamens 2;

ovaries glabrous;

haustorial synergids presumed absent;

styles 2, sometimes fused at the base, if free, close.

Lower glumes

1.3-3 mm, (1)3-5-veined;

upper glumes 1.3-2.5 mm, 3-5-veined;

calluses glabrous;

fertile lemmas 2.9-4.5 mm, straight, 3-7-veined, keels not winged, apices scabridulous;

paleas 2.3-3 mm;

anthers 1.3-1.5 mm, the length invariant within a spikelet.

Caryopses

1.9-2.2 mm, exposed at maturity.

Hila

basal, punctiform;

endosperm hard, without lipid;

starch grains simple;

embryos small or large relative to the caryopses;

epiblasts present;

scutellar cleft present;

mesocotyl internode present;

embryonic leaf margins overlapping, x = (11)12.

2n

= 24.

Chasmanthium laxum

Poaceae subfam. centothecoideae

Distribution
from FNA
AL; AR; DC; DE; FL; GA; KY; LA; MD; MO; MS; NC; NJ; NY; OK; PA; SC; TN; TX; VA
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[BONAP county map]
Discussion

Chasmanthium laxum is almost completely sympatric with C. sessiliflorum in the southeastern United States, growing in similar habitats but extending farther into sphagnous stream heads, pine flatwoods, and pine savannahs. Yates (1966b) reported seeing putative, naturally occurring hybrids between Chasmanthium ornithorhynchum and C. laxum along streams of the outer coastal plain of Mississippi and Louisiana. In general appearance, the hybrids resemble C. laxum, their most striking difference being the enlarged, sterile spikelets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The subfamily Centothecoideae is one of the subfamilies that cannot be characterized by a suite of morphological characteristics, but anatomical, micromorphological, and nucleic acid data all support its recognition. It is most abundant in warm-temperate woodlands and tropical forests. Clayton and Renvoize (1986) suggested that it was an offshoot of the Arundinoideae, but molecular data (Hilu et al. 1999; Grass Phylogeny Working Group 2001) argue for a sister group relationship with the Panicoideae. The treatment here, in which two tribes are recognized, follows that of the Grass Phylogeny Working Group (2001). Some of the genera, however, are as yet poorly known in terms of the characters used in making such decisions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 4-50 mm long, with 1-15 florets, the lowest florets sometimes sterile, the upper florets bisexual; disarticulation at the base of the florets or the base of the spikelets; leaves not pseudopetiolate; culms 35-150 cm tall; plants not reedlike
Centotheceae
1. Spikelets 1.2-1.8 mm long, with 2(3-4) florets, the lower florets sterile, the upper florets bisexual; disarticulation at the pedicel bases, subsequently below the spikelets; leaves pseudopetiolate; culms 150-400 cm tall; plants reedlike
Thysanolaeneae
Source FNA vol. 25, p. 346. FNA vol. 25, p. 343. Author: Grass Phylogeny Working Group;.
Parent taxa Poaceae > subfam. Centothecoideae > tribe Centotheceae > Chasmanthium Poaceae
Sibling taxa
C. latifolium, C. nitidum, C. ornithorhynchum, C. sessiliflorum
Synonyms Uniola laxa
Name authority (L.) H.O. Yates Soderstr.
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