Chaenactis thompsonii |
Chaenactis macrantha |
|
|---|---|---|
|
Thompson's pincushion |
big-head dusty maidens, bighead dustymaiden, large-flower chaenactis, Mohave pincushion, Mojave pincushion, showy dustymaidens |
|
| Habit | Perennials, 10–30 cm (not or scarcely cespitose, not matted); proximal indument thinning with age, grayish, arachnoid-sericeous to thinly lanuginose. | Plants 5–25(–35) cm; proximal indument grayish, arachnoid-sericeous to closely lanuginose (sometimes tardily glabrescent). |
| Stems | mostly 5–15+, ascending to erect. |
mostly 1–5; branches mainly proximal. |
| Leaves | mostly cauline, 2–5 cm; largest blades ± elliptic, ± plane, 1-pinnately lobed; lobes mostly 2–5 pairs, remote, ± plane. |
basal (withering) and cauline, 1.5–7 cm; largest blades ± elliptic to ovate, ± plane, not succulent, 1(–2)-pinnately lobed (± gland-dotted beneath indument); primary lobes mostly 2–5 pairs, ± remote, ultimate lobes ± plane. |
| Peduncles | ascending to erect, 2–5 cm. |
1.5–8 cm, arachnoid-sericeous to thinly lanuginose distally, not stipitate-glandular. |
| Involucres | ± obconic. |
± obconic to broadly cylindric. |
| Florets | corollas (nocturnal) white to pinkish or cream, 9–12(–15) mm (lengths 1.8–2.2 times cypselae; anthers ± included); peripheral corollas nocturnally spreading, actinomorphic, scarcely enlarged. |
|
| Corollas | 7–9 mm. |
|
| Phyllaries | longest (10–)12–15 mm; outer closely lanuginose, not stipitate-glandular, apices erect, ± rigid. |
longest 12–18 mm; outer arachnoid-sericeous to thinly lanuginose in fruit, not stipitate-glandular, apices ± squarrose, blunt, pliant. |
| Heads | mostly 1–3 per stem. |
(± radiant, nocturnally), mostly 1–5(–7) per stem (nodding in bud). |
| Cypselae | 7–9 mm (eglandular); pappi: longest scales 3.5–5 mm. |
5–6(–7) mm; pappi of 8 scales in 2, abruptly unequal series, longest scales 5–7 mm. |
| 2n | = 12. |
|
Chaenactis thompsonii |
Chaenactis macrantha |
|
| Phenology | Flowering Jun–Aug. | Flowering Mar–early Jul. |
| Habitat | Rocky or gravelly serpentine slopes, scree, talus, openings in or above conifer forests | Open, loose, light-colored, silty, usually calcareous or alkaline, desert soils, often covered by or mixed with gravel |
| Elevation | (900–)1200–2200 m [(3000–)3900–7200 ft] | 600–2200 m [2000–7200 ft] |
| Distribution |
WA
|
AZ; CA; ID; NV; OR; UT
|
| Discussion | Of conservation concern. Chaenactis thompsonii appears to be sister to C. evermannii; it is known from the mountains of central and northwestern Washington. The similar habits of C. thompsonii and C. ramosa (= C. douglasii var. douglasii) appear to result from convergent evolution in the distinctive habitat of their type localities (Wenatchee Mountains), not from a close genetic relationship as suggested by Cronquist. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though the derived floral features of Chaenactis macrantha obscure its relationships, it may represent a link between sect. Chaenactis (annuals; pappus scales in regular, often strongly reduced series) and sect. Macrocarphus (leaf blades gland-dotted). Resemblance of its heads, leaves, and indument to those of C. thompsonii and relatives is striking. It appears to form no natural hybrids, perhaps because of its nocturnal corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Name authority | Cronquist: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 5: 123, fig. [p. 125]. (1955) | D. C. Eaton: in S. Watson, Botany (Fortieth Parallel), 171, plate 18, figs. 1–5. (1871) |
| Source | FNA vol. 21, p. 407. | FNA vol. 21, p. 410. |
| Web links | ||