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Chaenactis thompsonii

Thompson's pincushion

common yellow chaenactis, yellow pincushion

Habit Perennials, 10–30 cm (not or scarcely cespitose, not matted); proximal indument thinning with age, grayish, arachnoid-sericeous to thinly lanuginose. Plants 6–60 cm; proximal indument grayish to whitish, arachnoid to densely lanuginose, or glabrescent.
Stems

mostly 5–15+, ascending to erect.

mostly 1–5(–12; sometimes ± horizontal);

branches proximal and, often, distal.

Leaves

mostly cauline, 2–5 cm;

largest blades ± elliptic, ± plane, 1-pinnately lobed;

lobes mostly 2–5 pairs, remote, ± plane.

basal (often withering) and cauline, 1–10 cm;

largest blades linear or ± elliptic, plane to 3-dimensional, succulent or not, (0–)1–2-pinnately lobed;

primary lobes 1–7 pairs, remote to ± congested, ultimate lobes ± plane, involute, twisted, and/or terete.

Peduncles

ascending to erect, 2–5 cm.

1–20(–30) cm, distally stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent.

Involucres

± obconic.

± hemispheric to obconic or broadly cylindric.

Florets

corollas bright to dark yellow, 4–8 mm (inner);

peripheral corollas spreading, ± zygomorphic, enlarged.

Corollas

7–9 mm.

Phyllaries

longest (10–)12–15 mm;

outer closely lanuginose, not stipitate-glandular, apices erect, ± rigid.

longest 4.5–10 mm;

outer stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent in fruit, apices erect, blunt, ± rigid.

Heads

mostly 1–3 per stem.

(± radiant) mostly 1–20+ per stem.

Cypselae

7–9 mm (eglandular);

pappi: longest scales 3.5–5 mm.

3–9 mm (± terete);

pappi of (1–)4 scales in 1 series, or of (5–)8 scales in 2, abruptly unequal series, longest scales (1–)2–8 mm.

2n

= 12.

Chaenactis thompsonii

Chaenactis glabriuscula

Phenology Flowering Jun–Aug.
Habitat Rocky or gravelly serpentine slopes, scree, talus, openings in or above conifer forests
Elevation (900–)1200–2200 m [(3000–)3900–7200 ft]
Distribution
map from FNA
WA
[WildflowerSearch map]
[BONAP county map]
map from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chaenactis thompsonii appears to be sister to C. evermannii; it is known from the mountains of central and northwestern Washington. The similar habits of C. thompsonii and C. ramosa (= C. douglasii var. douglasii) appear to result from convergent evolution in the distinctive habitat of their type localities (Wenatchee Mountains), not from a close genetic relationship as suggested by Cronquist.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

The diverse and intergrading forms here included in Chaenactis glabriuscula have been divided by P. Stockwell (1940) and subsequent workers into as many as four species and ten varieties. Chaenactis glabriuscula is known from the southern two-thirds of the Californian Floristic Province and adjacent desert edges. It has been reported in Massachusetts as a garden escape (variety unspecified); it is not expected to persist there outside cultivation.

Complete interfertility among the taxa recognized here as Chaenactis glabriuscula vars. glabriuscula, megacephala, and lanosa was demonstrated by P. Stockwell (1940). Intraspecific crosses involving C. glabriuscula var. orcuttiana were much less successful; C. glabriuscula var. heterocarpha was not tested. As noted by W. J. Hooker and G. A. W. Arnott ([1830–]1841) and D. W. Kyhos (1965), some forms of C. glabriuscula differ from C. stevioides or C. fremontii only in corolla color, which can be lost in older or poorly preserved specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis
Sibling taxa
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. xantiana
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Subordinate taxa
C. glabriuscula var. glabriuscula, C. glabriuscula var. heterocarpha, C. glabriuscula var. lanosa, C. glabriuscula var. megacephala, C. glabriuscula var. orcuttiana
Key
1. Pappi of (5–)8 scales in 2, abruptly unequal series
var. heterocarpha
1. Pappi of (1–)4(–5) scales in 1 series
→ 2
2. Proximal indument whitish, ± densely lanuginose; leaves ± basal (persistent)
var. lanosa
2. Proximal indument grayish, ± arachnoid, or glabrescent; leaves basal (withering) and cauline
→ 3
3. Largest leaf blades succulent, mostly 2-pinnately lobed; stems ± horizontal
var. orcuttiana
3. Largest leaf blades scarcely succulent, 1(–2)-pinnately lobed; stems ascending to erect
→ 4
4. Longest phyllaries 7–9 × 2–3 mm, outer glabrescent in fruit; pappi: lengths of longest scales ± 0.9(–1) times corollas
var. megacephala
4. Longest phyllaries 5–7 × 1–2 mm, outer (at least medially) ± arachnoid-sericeous and, often, ± stipitate-glandular in fruit; pappi: lengths of longest scales 0.4–0.7 times corollas
var. glabriuscula
Name authority Cronquist: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 5: 123, fig. [p. 125]. (1955) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 659. (1836)
Source FNA vol. 21, p. 407. Treatment author: James D. Morefield. FNA vol. 21, p. 411. Treatment author: James D. Morefield.
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