FNA: Chaenactis thompsonii vs. Chaenactis evermannii

	Chaenactis thompsonii	Chaenactis evermannii
	Thompson's pincushion	Evermann's pincushion
Habit	Perennials, 10–30 cm (not or scarcely cespitose, not matted); proximal indument thinning with age, grayish, arachnoid-sericeous to thinly lanuginose.	Perennials, mostly 6–12 cm (cespitose or ± matted); proximal indument thinning with age, grayish, mostly arachnoid-sericeous to thinly lanuginose.
Stems	mostly 5–15+, ascending to erect.	mostly 5–20+, ascending to erect.
Leaves	mostly cauline, 2–5 cm; largest blades ± elliptic, ± plane, 1-pinnately lobed; lobes mostly 2–5 pairs, remote, ± plane.	basal, 1–5 cm; largest blades broadly ± elliptic, ± plane, 1-pinnately lobed; lobes 2–5 pairs, remote, ± plane.
Peduncles	ascending to erect, 2–5 cm.	mostly ascending to erect, 5–10 cm.
Involucres	± obconic.	± obconic.
Corollas	7–9 mm.	5–6.5 mm.
Phyllaries	longest (10–)12–15 mm; outer closely lanuginose, not stipitate-glandular, apices erect, ± rigid.	longest 8–12 mm; outer predominantly closely lanuginose, sparsely, if at all, stipitate-glandular, apices erect, ± rigid.
Heads	mostly 1–3 per stem.	1(–3) per stem.
Cypselae	7–9 mm (eglandular); pappi: longest scales 3.5–5 mm.	5–6.5 mm (eglandular); pappi: longest scales 2.5–4.5 mm (lengths 0.4–0.8 times corollas).
2n		= 12.

	Chaenactis thompsonii	Chaenactis evermannii
Phenology	Flowering Jun–Aug.	Flowering Jun–Aug.
Habitat	Rocky or gravelly serpentine slopes, scree, talus, openings in or above conifer forests	Subalpine, usually decomposing granitic sand or gravel slopes, ridges, scree, talus, openings in or above conifer forests
Elevation	(900–)1200–2200 m ((3000–)3900–7200 ft)	1200–3000 m (3900–9800 ft)
Distribution	WA [WildflowerSearch map] [BONAP county map]	ID [BONAP county map]
Discussion	Of conservation concern. Chaenactis thompsonii appears to be sister to C. evermannii; it is known from the mountains of central and northwestern Washington. The similar habits of C. thompsonii and C. ramosa (= C. douglasii var. douglasii) appear to result from convergent evolution in the distinctive habitat of their type localities (Wenatchee Mountains), not from a close genetic relationship as suggested by Cronquist. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Chaenactis evermannii is known from mountains of central Idaho. Reports of it from Washington and California/Nevada (P. Stockwell 1940, some as C. nevadensis var. mainsiana) were based on specimens of C. thompsonii and C. alpigena, respectively; all three species are closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 21, p. 407.	FNA vol. 21, p. 408.
Parent taxa	Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus	Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus
Sibling taxa	C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. xantiana	C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Synonyms		C. nevadensis var. mainsiana
Name authority	Cronquist: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 5: 123, fig. [p. 125]. (1955)	Greene: Leafl. Bot. Observ. Crit. 2: 224. (1912)
Web links	Local floras: WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Chaenactis evermannii

Thompson's pincushion

Evermann's pincushion

Habit

Perennials, 10–30 cm (not or scarcely cespitose, not matted); proximal indument thinning with age, grayish, arachnoid-sericeous to thinly lanuginose.

Perennials, mostly 6–12 cm (cespitose or ± matted); proximal indument thinning with age, grayish, mostly arachnoid-sericeous to thinly lanuginose.

Stems

mostly 5–15+, ascending to erect.

mostly 5–20+, ascending to erect.

Leaves

mostly cauline, 2–5 cm;

largest blades ± elliptic, ± plane, 1-pinnately lobed;

lobes mostly 2–5 pairs, remote, ± plane.

basal, 1–5 cm;

largest blades broadly ± elliptic, ± plane, 1-pinnately lobed;

lobes 2–5 pairs, remote, ± plane.

Peduncles

ascending to erect, 2–5 cm.

mostly ascending to erect, 5–10 cm.

Involucres

± obconic.

Corollas

7–9 mm.

5–6.5 mm.

Phyllaries

longest (10–)12–15 mm;

outer closely lanuginose, not stipitate-glandular, apices erect, ± rigid.

longest 8–12 mm;

outer predominantly closely lanuginose, sparsely, if at all, stipitate-glandular, apices erect, ± rigid.

Heads

mostly 1–3 per stem.

1(–3) per stem.

Cypselae

7–9 mm (eglandular);

pappi: longest scales 3.5–5 mm.

5–6.5 mm (eglandular);

pappi: longest scales 2.5–4.5 mm (lengths 0.4–0.8 times corollas).

= 12.

Chaenactis evermannii

Phenology

Flowering Jun–Aug.

Habitat

Rocky or gravelly serpentine slopes, scree, talus, openings in or above conifer forests

Subalpine, usually decomposing granitic sand or gravel slopes, ridges, scree, talus, openings in or above conifer forests

Elevation

(900–)1200–2200 m ((3000–)3900–7200 ft)

1200–3000 m (3900–9800 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

Discussion

Of conservation concern.

Chaenactis thompsonii appears to be sister to C. evermannii; it is known from the mountains of central and northwestern Washington. The similar habits of C. thompsonii and C. ramosa (= C. douglasii var. douglasii) appear to result from convergent evolution in the distinctive habitat of their type localities (Wenatchee Mountains), not from a close genetic relationship as suggested by Cronquist.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chaenactis evermannii is known from mountains of central Idaho. Reports of it from Washington and California/Nevada (P. Stockwell 1940, some as C. nevadensis var. mainsiana) were based on specimens of C. thompsonii and C. alpigena, respectively; all three species are closely related.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 21, p. 407.

FNA vol. 21, p. 408.

Parent taxa

Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus

Sibling taxa

C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. xantiana

C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana

Synonyms

C. nevadensis var. mainsiana

Name authority

Cronquist: in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 5: 123, fig. [p. 125]. (1955)

Greene: Leafl. Bot. Observ. Crit. 2: 224. (1912)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, UW Burke Herbarium

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Chaenactis thompsonii

Chaenactis evermannii

Chaenactis thompsonii

Chaenactis evermannii