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broad-flower chaenactis, desert or Esteve or broad-flower pincushion, desert pincushion, Esteve pincushion, Esteve's pincushion, Steve's dustymaiden

santolina chaenactis, santolina pincushion

Habit Plants 5–30(–45) cm; proximal indument grayish, ± arachnoid-sericeous (tardily glabrescent except around nodes). Perennials, 10–25(–35) cm (cespitose or ± matted); proximal indument thinning with age, whitish to grayish, lanuginose.
Stems

1–12 (sometimes decumbent);

branches proximal and/or distal.

mostly 5–15+, erect to ± spreading.

Leaves

basal (usually withering) and ± cauline, 1–8(–10) cm;

largest blades ± elliptic, ± 3-dimensional, usually not succulent, mostly 1–2-pinnately lobed;

primary lobes 4–8 pairs, remote or ± congested, ultimate lobes ± involute and/or twisted.

basal, (1–)3–11 cm;

largest blades linear-cylindric to ± fusiform, 3-dimensional, 1–2-pinnately lobed;

primary lobes (7–)10–18+ pairs, ± imbricate, ultimate lobes ± involute, twisted.

Peduncles

1–5(–10) cm, usually stipitate-glandular distally and, often, ± arachnoid.

mostly ascending to erect, mostly 8–25 cm.

Involucres

± hemispheric to obconic (bases green, rounded in fruit).

obconic to ± cylindric.

Florets

corollas white to pinkish, cream, or pale yellow, 4.5–6.5 mm (inner);

peripheral corollas spreading, zygomorphic, enlarged.

Corollas

5–7 mm.

Phyllaries

longest 5.5–8(–10) mm;

outer stipitate-glandular and/or ± arachnoid in fruit, apices erect, blunt, ± rigid.

longest 8–13 mm;

outer evidently stipitate-glandular and, sometimes, ± arachnoid, apices erect, ± rigid.

Heads

(± radiant) mostly 3–20+ per stem.

1(–3) per stem.

Cypselae

(3–)4–6.5 mm;

pappi of (1–)4(–5) scales, usually in 1 series, rarely with partial outer, abruptly unequal series, longest scales 1.5–6 mm, lengths mostly 0.3–0.9 times corollas (apices hidden among corollas at flowering).

4–6 mm;

pappi: longest scales 3–4.5 mm.

2n

= 10.

= 12.

Chaenactis stevioides

Chaenactis santolinoides

Phenology Flowering Feb–Jun. Flowering (Mar–)May–Jul.
Habitat Open, arid or semiarid, sandy or gravelly slopes and flats, shrublands Exposed sandy to rocky summits, ridges, scree, talus, openings in or above conifer forests, sometimes road cuts or other recent disturbances
Elevation -30–2100(–2300) m (-100–6900(–7500) ft) (1100–)1500–2800 m ((3600–)4900–9200 ft)
Distribution
from FNA
AZ; CA; CO; ID; NM; NV; OR; UT; WY; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chaenactis stevioides is found throughout the southwestern deserts; it is among the most abundant spring wildflowers in the higher Mojave Desert and southern Great Basin. It also extends seaward into west-central California. It has been reported in New York as a garden escape; it is not expected to persist there outside cultivation.

Chaenactis stevioides varies in more or less concentric zones. Plants from the core zone (centered on the Great Basin and Mojave Desert) typically have pappi and phyllaries relatively short and phyllaries predominantly stipitate-glandular (var. brachypappa). Surrounding this zone to the southwest, southeast, and northeast are plants with pappi and phyllaries relatively long and phyllaries evidently or predominantly lanuginose (var. stevioides). Scattered on the periphery in central Arizona, Baja California, and west-central and southwestern California (where hybrids may be involved; see sectional discussion) are mesophytic forms with relatively long and/or broad leaf divisions, corollas varying from white to pale yellow, and pappi and phyllaries like those of var. brachypappa (var. thornberi, C. gillespiei). An unnamed form with leaves arachnoid but otherwise like C. fremontii occurs around sand dunes in the Mojave Desert. Chaenactis furcata and C. latifolia are forms possibly influenced by C. fremontii genes, unusual substrates, or pathogens. Traits of all the above taxa are inconsistent within populations, and/or recurrent or recombinant elsewhere in the range of C. stevioides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chaenactis santolinoides is known from the southern Sierra Nevada and Transverse Ranges. It is sometimes cultivated in rock gardens and may be found beyond its native range. It may be relatively recently derived from an isolated segment of C. douglasii var. alpina. The morphology of C. panamintensis (here assigned to C. douglasii var. alpina) suggests past convergence toward, or genetic influence from, C. santolinoides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 413. FNA vol. 21, p. 407.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus
Sibling taxa
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. suffrutescens, C. thompsonii, C. xantiana
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Synonyms C. furcata, C. gillespiei, C. latifolia, C. mexicana, C. stevioides var. brachypappa, C. stevioides var. thornberi
Name authority Hooker & Arnott: Bot. Beechey Voy., 353. (1839) Greene: Bull. Torrey Bot. Club 9: 17. (1882)
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