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broad-flower chaenactis, desert or Esteve or broad-flower pincushion, desert pincushion, Esteve pincushion, Esteve's pincushion, Steve's dustymaiden

Parish chaenactis, Parish's chaenactis

Habit Plants 5–30(–45) cm; proximal indument grayish, ± arachnoid-sericeous (tardily glabrescent except around nodes). Subshrubs, (10–)20–40(–60) cm (not cespitose or matted); proximal indument (especially of stems) persistent, whitish, densely lanuginose or pannose.
Stems

1–12 (sometimes decumbent);

branches proximal and/or distal.

mostly 5–15+, erect.

Leaves

basal (usually withering) and ± cauline, 1–8(–10) cm;

largest blades ± elliptic, ± 3-dimensional, usually not succulent, mostly 1–2-pinnately lobed;

primary lobes 4–8 pairs, remote or ± congested, ultimate lobes ± involute and/or twisted.

mostly cauline, (1–)2–5 cm;

largest blades lance-ovate or deltate, ± plane, 1-pinnately lobed;

lobes mostly 2–5 pairs, remote, ± plane.

Peduncles

1–5(–10) cm, usually stipitate-glandular distally and, often, ± arachnoid.

ascending to erect, 2–8(–20) cm.

Involucres

± hemispheric to obconic (bases green, rounded in fruit).

± obconic.

Florets

corollas white to pinkish, cream, or pale yellow, 4.5–6.5 mm (inner);

peripheral corollas spreading, zygomorphic, enlarged.

Corollas

7–8.5 mm.

Phyllaries

longest 5.5–8(–10) mm;

outer stipitate-glandular and/or ± arachnoid in fruit, apices erect, blunt, ± rigid.

longest 10–13 mm;

outer predominantly arachnoid to closely lanuginose (sparsely, if at all, stipitate-glandular), apices ± squarrose, pliant.

Heads

(± radiant) mostly 3–20+ per stem.

mostly 1–3 per stem.

Cypselae

(3–)4–6.5 mm;

pappi of (1–)4(–5) scales, usually in 1 series, rarely with partial outer, abruptly unequal series, longest scales 1.5–6 mm, lengths mostly 0.3–0.9 times corollas (apices hidden among corollas at flowering).

4–7 mm;

pappi: longest scales 6–8 mm.

2n

= 10.

= 12.

Chaenactis stevioides

Chaenactis parishii

Phenology Flowering Feb–Jun. Flowering May–Jul.
Habitat Open, arid or semiarid, sandy or gravelly slopes and flats, shrublands Open rocky to sandy soils in low montane chaparral
Elevation -30–2100(–2300) m (-100–6900(–7500) ft) 1300–2500 m (4300–8200 ft)
Distribution
from FNA
AZ; CA; CO; ID; NM; NV; OR; UT; WY; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[BONAP county map]
Discussion

Chaenactis stevioides is found throughout the southwestern deserts; it is among the most abundant spring wildflowers in the higher Mojave Desert and southern Great Basin. It also extends seaward into west-central California. It has been reported in New York as a garden escape; it is not expected to persist there outside cultivation.

Chaenactis stevioides varies in more or less concentric zones. Plants from the core zone (centered on the Great Basin and Mojave Desert) typically have pappi and phyllaries relatively short and phyllaries predominantly stipitate-glandular (var. brachypappa). Surrounding this zone to the southwest, southeast, and northeast are plants with pappi and phyllaries relatively long and phyllaries evidently or predominantly lanuginose (var. stevioides). Scattered on the periphery in central Arizona, Baja California, and west-central and southwestern California (where hybrids may be involved; see sectional discussion) are mesophytic forms with relatively long and/or broad leaf divisions, corollas varying from white to pale yellow, and pappi and phyllaries like those of var. brachypappa (var. thornberi, C. gillespiei). An unnamed form with leaves arachnoid but otherwise like C. fremontii occurs around sand dunes in the Mojave Desert. Chaenactis furcata and C. latifolia are forms possibly influenced by C. fremontii genes, unusual substrates, or pathogens. Traits of all the above taxa are inconsistent within populations, and/or recurrent or recombinant elsewhere in the range of C. stevioides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chaenactis parishii is sometimes cultivated in rock gardens. It is known from small, isolated populations in the higher Peninsular Ranges of Riverside and San Diego counties and adjacent Baja California. Chaenactis parishii and C. suffrutescens form a species pair well marked by the (usually) subshrubby habit, proximal indument persistent, white, felty, heads relatively large, and largest leaf blades lance-ovate to deltate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 413. FNA vol. 21, p. 404.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus
Sibling taxa
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. suffrutescens, C. thompsonii, C. xantiana
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Synonyms C. furcata, C. gillespiei, C. latifolia, C. mexicana, C. stevioides var. brachypappa, C. stevioides var. thornberi
Name authority Hooker & Arnott: Bot. Beechey Voy., 353. (1839) A. Gray: Proc. Amer. Acad. Arts 20: 299. (1885)
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