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common yellow chaenactis, yellow pincushion

Parish chaenactis, Parish's chaenactis

Habit Plants 6–60 cm; proximal indument grayish to whitish, arachnoid to densely lanuginose, or glabrescent. Subshrubs, (10–)20–40(–60) cm (not cespitose or matted); proximal indument (especially of stems) persistent, whitish, densely lanuginose or pannose.
Stems

mostly 1–5(–12; sometimes ± horizontal);

branches proximal and, often, distal.

mostly 5–15+, erect.

Leaves

basal (often withering) and cauline, 1–10 cm;

largest blades linear or ± elliptic, plane to 3-dimensional, succulent or not, (0–)1–2-pinnately lobed;

primary lobes 1–7 pairs, remote to ± congested, ultimate lobes ± plane, involute, twisted, and/or terete.

mostly cauline, (1–)2–5 cm;

largest blades lance-ovate or deltate, ± plane, 1-pinnately lobed;

lobes mostly 2–5 pairs, remote, ± plane.

Peduncles

1–20(–30) cm, distally stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent.

ascending to erect, 2–8(–20) cm.

Involucres

± hemispheric to obconic or broadly cylindric.

± obconic.

Florets

corollas bright to dark yellow, 4–8 mm (inner);

peripheral corollas spreading, ± zygomorphic, enlarged.

Corollas

7–8.5 mm.

Phyllaries

longest 4.5–10 mm;

outer stipitate-glandular, ± villous, arachnoid-sericeous, lanuginose, and/or glabrescent in fruit, apices erect, blunt, ± rigid.

longest 10–13 mm;

outer predominantly arachnoid to closely lanuginose (sparsely, if at all, stipitate-glandular), apices ± squarrose, pliant.

Heads

(± radiant) mostly 1–20+ per stem.

mostly 1–3 per stem.

Cypselae

3–9 mm (± terete);

pappi of (1–)4 scales in 1 series, or of (5–)8 scales in 2, abruptly unequal series, longest scales (1–)2–8 mm.

4–7 mm;

pappi: longest scales 6–8 mm.

2n

= 12.

= 12.

Chaenactis glabriuscula

Chaenactis parishii

Phenology Flowering May–Jul.
Habitat Open rocky to sandy soils in low montane chaparral
Elevation 1300–2500 m (4300–8200 ft)
Distribution
from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

The diverse and intergrading forms here included in Chaenactis glabriuscula have been divided by P. Stockwell (1940) and subsequent workers into as many as four species and ten varieties. Chaenactis glabriuscula is known from the southern two-thirds of the Californian Floristic Province and adjacent desert edges. It has been reported in Massachusetts as a garden escape (variety unspecified); it is not expected to persist there outside cultivation.

Complete interfertility among the taxa recognized here as Chaenactis glabriuscula vars. glabriuscula, megacephala, and lanosa was demonstrated by P. Stockwell (1940). Intraspecific crosses involving C. glabriuscula var. orcuttiana were much less successful; C. glabriuscula var. heterocarpha was not tested. As noted by W. J. Hooker and G. A. W. Arnott ([1830–]1841) and D. W. Kyhos (1965), some forms of C. glabriuscula differ from C. stevioides or C. fremontii only in corolla color, which can be lost in older or poorly preserved specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chaenactis parishii is sometimes cultivated in rock gardens. It is known from small, isolated populations in the higher Peninsular Ranges of Riverside and San Diego counties and adjacent Baja California. Chaenactis parishii and C. suffrutescens form a species pair well marked by the (usually) subshrubby habit, proximal indument persistent, white, felty, heads relatively large, and largest leaf blades lance-ovate to deltate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pappi of (5–)8 scales in 2, abruptly unequal series
var. heterocarpha
1. Pappi of (1–)4(–5) scales in 1 series
→ 2
2. Proximal indument whitish, ± densely lanuginose; leaves ± basal (persistent)
var. lanosa
2. Proximal indument grayish, ± arachnoid, or glabrescent; leaves basal (withering) and cauline
→ 3
3. Largest leaf blades succulent, mostly 2-pinnately lobed; stems ± horizontal
var. orcuttiana
3. Largest leaf blades scarcely succulent, 1(–2)-pinnately lobed; stems ascending to erect
→ 4
4. Longest phyllaries 7–9 × 2–3 mm, outer glabrescent in fruit; pappi: lengths of longest scales ± 0.9(–1) times corollas
var. megacephala
4. Longest phyllaries 5–7 × 1–2 mm, outer (at least medially) ± arachnoid-sericeous and, often, ± stipitate-glandular in fruit; pappi: lengths of longest scales 0.4–0.7 times corollas
var. glabriuscula
Source FNA vol. 21, p. 411. FNA vol. 21, p. 404.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus
Sibling taxa
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Subordinate taxa
C. glabriuscula var. glabriuscula, C. glabriuscula var. heterocarpha, C. glabriuscula var. lanosa, C. glabriuscula var. megacephala, C. glabriuscula var. orcuttiana
Name authority de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 659. (1836) A. Gray: Proc. Amer. Acad. Arts 20: 299. (1885)
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