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chaenactis, Douglas' dusty maidens, Douglas' dustymaiden, hoary chaenactis, hoary false-yarrow, hoary pincushion

Cusick's pincushion, morning brides

Habit Biennials or perennials, (2–)5–50(–60) cm (rarely slightly woody or flowering first year, sometimes cespitose or ± matted); proximal indument thinning with age, grayish, mostly arachnoid-sericeous to thinly lanuginose. Plants 3.5–10(–15) cm; proximal indument grayish, sparsely arachnoid, glabrescent.
Stems

1–25+, erect to spreading.

mostly 1–5;

branches mainly proximal.

Leaves

basal, or basal (sometimes withering) and ± cauline, (1–)2–12(–15) cm;

largest blades ± elliptic or slightly lanceolate to ovate, ± 3-dimensional, usually 2-pinnately lobed;

primary lobes (4–)5–9(–12) pairs, ± congested, scarcely imbricate, ultimate lobes ± involute and/or twisted.

basal (withering) and cauline (not notably smaller or sparser distally), 1–4 cm;

largest blades ± oblanceolate, plane, ± succulent, not lobed (margins entire or distally ± crenate).

Peduncles

mostly ascending to erect, 1–10 cm.

0.5–2.5 cm, glabrescent distally (sparsely arachnoid early; bracts 1–2, leaflike, surpassing heads).

Involucres

obconic to ± hemispheric.

± hemispheric to campanulate.

Florets

corollas (diurnal) white to pinkish, 3–7 mm (± equaling cypselae; anthers exserted);

peripheral corollas ± erect, actinomorphic, scarcely enlarged.

Corollas

5–8 mm.

Phyllaries

longest 9–15(–17) mm;

outer usually stipitate-glandular (sometimes sparsely or obscurely, rarely eglandular) and, often, arachnoid to lanuginose and, sometimes, sparsely villous, apices usually ± squarrose, pliant.

longest 6–9(–10) mm (surpassed by florets);

outer (uniformly) sparsely arachnoid to glabrescent in fruit, not stipitate-glandular (inner apically brownish-villosulous), apices usually erect, blunt, ± rigid.

Heads

1–25+ per stem.

mostly 1–5(–12) per stem.

Cypselae

5–8 mm (usually sparsely glandular amidst other indument);

pappi: longest scales 3–6 mm.

4–6 mm;

pappi of (8–)10–14 scales in 2–3 gradually unequal series, longest scales 1.5–3.5 mm.

2n

= 12.

Chaenactis douglasii

Chaenactis cusickii

Phenology Flowering late Apr–Jun.
Habitat Light-colored shrink-swell clay soils from volcanic ash and tuff, semiarid shrublands
Elevation 700–1300(–1800) m (2300–4300(–5900) ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; ND; NM; NV; OR; SD; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Chaenactis douglasii is widespread and variable (see discussion under var. douglasii).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chaenactis cusickii is known from Malheur County, Oregon, and adjacent Owyhee and Canyon counties, Idaho. Its relationship to other species is obscure.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves basal (sometimes withering) and ± cauline; plants not or scarcely cespitose, not matted; stems usually 1–5(–12); heads (1–)2–25+ per stem
var. douglasii
1. Leaves strictly basal; plants cespitose or ± matted; stems (1–)10–25+; heads 1(–2) per stem
var. alpina
Source FNA vol. 21, p. 405. FNA vol. 21, p. 410.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Macrocarphus Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis
Sibling taxa
C. alpigena, C. artemisiifolia, C. carphoclinia, C. cusickii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
C. alpigena, C. artemisiifolia, C. carphoclinia, C. douglasii, C. evermannii, C. fremontii, C. glabriuscula, C. macrantha, C. nevadensis, C. nevii, C. parishii, C. santolinoides, C. stevioides, C. suffrutescens, C. thompsonii, C. xantiana
Subordinate taxa
C. douglasii var. alpina, C. douglasii var. douglasii
Synonyms Hymenopappus douglasii, Macrocarphus douglasii
Name authority (Hooker) Hooker & Arnott: Bot. Beechey Voy., 354. (1839) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. ed. 2. 1(2): 452. (1886)
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