Chaenactis artemisiifolia |
Chaenactis macrantha |
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artemisia leaf chaenactis, white pincushion |
big-head dusty maidens, bighead dustymaiden, large-flower chaenactis, Mohave pincushion, Mojave pincushion, showy dustymaidens |
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Habit | Plants (15–)25–90(–200) cm. | Plants 5–25(–35) cm; proximal indument grayish, arachnoid-sericeous to closely lanuginose (sometimes tardily glabrescent). |
Stems | mostly 1–5; branches mainly proximal. |
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Leaves | basal (withering) and cauline, 3–15(–20) cm; largest blades ± plane, not succulent; primary lobes mostly 5–10 pairs, ultimate lobes ± crowded, antrorse, lanceolate to elliptic, plane. |
basal (withering) and cauline, 1.5–7 cm; largest blades ± elliptic to ovate, ± plane, not succulent, 1(–2)-pinnately lobed (± gland-dotted beneath indument); primary lobes mostly 2–5 pairs, ± remote, ultimate lobes ± plane. |
Peduncles | 1.5–6 cm. |
1.5–8 cm, arachnoid-sericeous to thinly lanuginose distally, not stipitate-glandular. |
Involucres | ± hemispheric, mostly 10–15 mm diam. |
± obconic to broadly cylindric. |
Receptacles | paleae 0. |
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Florets | corollas (nocturnal) white to pinkish or cream, 9–12(–15) mm (lengths 1.8–2.2 times cypselae; anthers ± included); peripheral corollas nocturnally spreading, actinomorphic, scarcely enlarged. |
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Corollas | 5–7 mm. |
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Phyllaries | longest 7–10(–12) mm, ± densely villous, not or sparsely glandular; apices (all) erect, ± green, acute or scarcely acuminate, not aristate, ± plane. |
longest 12–18 mm; outer arachnoid-sericeous to thinly lanuginose in fruit, not stipitate-glandular, apices ± squarrose, blunt, pliant. |
Heads | (± radiant, nocturnally), mostly 1–5(–7) per stem (nodding in bud). |
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Cypselae | compressed, 4–7 mm; pappi 0 or coroniform (of ± 10 scales, longest 0.1–0.5 mm). |
5–6(–7) mm; pappi of 8 scales in 2, abruptly unequal series, longest scales 5–7 mm. |
2n | = 16. |
= 12. |
Chaenactis artemisiifolia |
Chaenactis macrantha |
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Phenology | Flowering Apr–early Jul. | Flowering Mar–early Jul. |
Habitat | Dry canyons, open slopes, often over granitoid rocks, locally abundant in chaparral burns or other recovering disturbances | Open, loose, light-colored, silty, usually calcareous or alkaline, desert soils, often covered by or mixed with gravel |
Elevation | 80–1600 m (300–5200 ft) | 600–2200 m (2000–7200 ft) |
Distribution |
CA; Mexico (Baja California)
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AZ; CA; ID; NV; OR; UT
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Discussion | In the flora area, Chaenactis artemisiifolia is known from the Transverse and Peninsular ranges and seaward valleys of southwestern California. It is fire-adapted; its germination is significantly enhanced by exposure to biomass smoke (J. E. Keeley and C. J. Fotheringham 1998). Chaenactis lacera Greene, the eighteenth species of the genus, is known from coastal portions (including islands) of the western Vizcaíno Desert in Baja California and Baja California Sur, Mexico. Forms of C. artemisiifolia sometimes resemble C. lacera in coastal southern California (P. Stockwell 1940), where C. lacera could eventually be introduced. Besides the key characteristics above, C. lacera differs from C. artemisiifolia by its largest leaf blades broadly ± elliptic, 2–3-pinnately lobed, ultimate lobes remote, recurved to retrorse, ± linear, involute (leaf blades appearing ± skeletal). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though the derived floral features of Chaenactis macrantha obscure its relationships, it may represent a link between sect. Chaenactis (annuals; pappus scales in regular, often strongly reduced series) and sect. Macrocarphus (leaf blades gland-dotted). Resemblance of its heads, leaves, and indument to those of C. thompsonii and relatives is striking. It appears to form no natural hybrids, perhaps because of its nocturnal corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 402. | FNA vol. 21, p. 410. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Acarphaea | Asteraceae > tribe Heliantheae > subtribe Chaenactidinae > Chaenactis > sect. Chaenactis |
Sibling taxa | ||
Synonyms | Acarphaea artemisiifolia | |
Name authority | (Harvey & A. Gray) A. Gray: Proc. Amer. Acad. Arts 10: 74. (1874) | D. C. Eaton: in S. Watson, Botany (Fortieth Parallel), 171, plate 18, figs. 1–5. (1871) |
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