Ceratodon purpureus |
Ceratodon heterophyllus |
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ceratodon moss, fire-moss |
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Habit | Plants in open to dense tufts, turfs, or mats, green, dark green, brownish green, light green or yellow-green, usually darker proximally, often tinged reddish brown or purple. | Plants in dense tufts or mats, green to dark green to brownish green. | ||||||||
Stems | (0.2–)1–3(–4) cm. |
0.3–1 cm. |
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Leaves | crowded, erect-patent to contorted or somewhat crisped, rarely straight when dry, lanceolate, ovate-lanceolate, or triangular-lanceolate, 0.35–2.8 mm, margins recurved to near apex or rarely plane, irregularly serrate to uneven or smooth distally, apices acute to short-acuminate or, rarely, obtuse; costa strong, sub-percurrent to excurrent, sometimes as a long, smooth awn, medial laminal cells (6.5–)8–12(–14) µm, cell walls even, usually of medium thickness, often somewhat thicker and rounded at the cell angles. |
loosely imbricate, somewhat contorted when dry, ovate to ovate-lanceolate, concave and often somewhat cucullate, (0.35–)0.5–0.9(–1.5) mm, margins plane to weakly recurved, often only at mid-leaf, and entire to, rarely, weakly toothed distally, apices obtuse; most costae ending before apex; medial laminal cells (9–)12–16(–22) µm, thin-walled to somewhat thickened. |
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Seta | 1–3(–4) cm, various shades of red, orange, or yellow. |
(0.9–)1.5(–2.5) cm, red. |
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Capsule | oblong to long-cylindric, (1–)2–2.5(–3) mm, smooth to strongly sulcate when dry; free to united at their nodes, finely papillose to spinulose-papillose, dark red and bordered to completely pale and absent borders. |
ovate-cylindric to, occasionally, ovate, (0.9–)1–1.2(–1.5) mm, strongly sulcate when dry; peristome segments free at their nodes, finely papillose, dark red with pale borders to, rarely, completely pale. |
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Spores | (10–)11–14(–17) µm. |
(18–)19–21(–22) µm. |
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Ceratodon purpureus |
Ceratodon heterophyllus |
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Phenology | Capsules mature early summer–late fall. | |||||||||
Habitat | Sandy, usually moist soil, Arctic habitats | |||||||||
Elevation | low elevations (0-200 m) (low elevations (0-700 ft)) | |||||||||
Distribution |
Nearly worldwide
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AK; NT; YT; Greenland; Eurasia |
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Discussion | Subspecies 4 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In Northwest Territories, Ceratodon heterophyllus is found only on Prince Patrick Island. Ceratodon heterophyllus has also been treated as C. purpureus forma heterophyllus (Kindberg) Britton, and R. R. Ireland (1980) considered it to be a variety of C. purpureus, but J. S. Burley and N. M. Pritchard (1990) provided ample evidence that it is a distinct species. The broadly ovate leaves of C. heterophyllus grade into some alpine variations of C. purpureus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 446. | FNA vol. 27, p. 446. | ||||||||
Parent taxa | Ditrichaceae > Ceratodon | Ditrichaceae > Ceratodon | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Dicranum purpureum, C. purpurascens, C. purpureus var. purpurascens, C. purpureus var. xanthopus | C. purpureus var. obtusifolius, C. purpureus var. rotundifolius | ||||||||
Name authority | (Hedwig) Bridel: Bryol. Univ. 1:480. (1826) | Kindberg: Ottawa Naturalist 5: 179. (1892) | ||||||||
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