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céraiste tomenteux, dusty miller, snow-in-summer, snow-in-summer chickweed, tomentose chickweed

large field mouse-ear chickweed

Habit Plants perennial, mat-forming, rhizomatous. Plants perennial, usually clumped, taproot present, sometimes with short rhizomes.
Stems

flowering stems ascending, branched, 15–40 cm; nonflowering stems prostrate proximally, rooting readily, pubescence dense, white-tomentose, eglandular; small axillary tufts of leaves often present.

flowering shoots ascending, often decumbent at base, branched, (15–)25–35(–40) cm, softly pubescent to subglabrous, hairs spreading, straight, concentrated in longitudinal lines toward base, glandular and eglandular; nonflowering leafy shoots often present, straggling, 5–15 cm; small axillary tufts of leaves present.

Leaves

not marcescent, sessile;

blade linear to linear-lanceolate or linear-oblong, 10–60 × 2–8 mm, apex ± obtuse, pubescence dense, whitish-tomentose, eglandular on both surfaces.

sessile;

cauline with largest at mid stem, smaller distally and proximally, blade narrowly oblong, linear-lanceolate to lanceolate or ovate, 20–45 × 3–11 mm, apex usually obtuse, softly ciliate-pubescent on both surfaces, more so on margins and abaxial midrib, rarely subglabrous, sometimes densely villose;

proximal leaf blades often oblanceolate, apex ± obtuse.

Inflorescences

lax, 3–13-flowered cymes;

bracts lanceolate, margins scarious, pubescent.

lax, 5–20-flowered cymes, pubescent, glandular, not viscid;

bracts lanceolate, proximal foliaceous, distal smaller, with scarious margins, glandular-pubescent.

Pedicels

ascending, straight, 10–40 mm, 2–7 times as long as sepals, white-tomentose.

curved immediately below capsule, slender, 10–24 mm, ca. 2 times as long as sepals, pubescence spreading, glandular.

Flowers

12–20 mm diam.;

sepals narrowly lanceolate-elliptic, 5–7 mm, margins narrow, often scarious, apex acute, white-tomentose;

petals obtriangular, 10–18 mm, 2–2.5 times as long as sepals, apex 2-fid;

stamens 10;

styles 5.

sepals lanceolate-elliptic, 5–8 mm, margins narrow, apex acute, densely and softly glandular-pubescent;

petals obovate, 10–15 mm, 2 times as long as sepals, apex deeply 2-fid;

stamens 10;

anthers 0.9–1.1 mm;

styles 5.

Capsules

cylindric, slightly curved, 10–15 mm, 1.5–2 times as long as sepals;

teeth 10, erect, margins convolute.

cylindric, slightly curved, slender, 10–14 × 3–4 mm, 2–2.3 times as long as sepals;

teeth 10, erect, margins convolute.

Seeds

brown, ca. 1.5 mm, round tubercles on margins, faces shallowly rugose;

testa not inflated.

brown, 0.8–1.2 mm diam., tuberculate;

testa not inflated, tightly enclosing seed.

2n

= 72.

= 72.

Cerastium tomentosum

Cerastium velutinum

Phenology Flowering spring.
Habitat A commonly grown rock-garden and wall plant, often escaping onto roadsides, riverbanks, old fields
Elevation 0-400 m (0-1300 ft)
Distribution
from FNA
ME; MI; MT; NC; NE; NY; OH; OR; PA; WA; WI; WY; BC; MB; NB; NF; NS; ON; PE; QC; se Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
DC; DE; IA; IL; IN; KY; MA; MD; MI; MN; MO; NJ; NY; OH; PA; TN; VA; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Cerastium tomentosum hybridizes readily with the introduced C. arvense subsp. arvense (J. K. Morton 1973).

North American reports of Cerastium biebersteinii de Candolle all appear to be referable to C. tomentosum. The two species are very similar, but C. biebersteinii has flat capsule teeth and is diploid (2n = 36); see M. K. Khalaf and C. A. Stace (2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Until recently Cerastium velutinum has been included in C. arvense as a variety or subspecies. However, morphological characters and chromosome number (diploid in the native forms of C. arvense, tetraploid in C. velutinum) distinguish the two taxa. They are not interfertile. Crosses between them can be produced with difficulty but they are completely sterile. None are known to occur naturally.

An apparent hybrid between Cerastium velutinum and C. tometosum has been collected on the south shore of Lake Erie. It was completely sterile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaves narrowly oblong to linear-lanceolate or lanceolate, not overlapping to obscure stem, not marcescent, softly pubescent to subglabrous
var. velutinum
1. Cauline leaves lanceolate to ovate, dense and overlapping, obscuring stem on sterile shoots, often marcescent, densely villous
var. villosissimum
Source FNA vol. 5, p. 91. FNA vol. 5, p. 91.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Cerastium Caryophyllaceae > subfam. Alsinoideae > Cerastium
Sibling taxa
C. aleuticum, C. alpinum, C. arcticum, C. arvense, C. axillare, C. beeringianum, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. fontanum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. velutinum, C. viride
C. aleuticum, C. alpinum, C. arcticum, C. arvense, C. axillare, C. beeringianum, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. fontanum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. tomentosum, C. viride
Subordinate taxa
C. velutinum var. velutinum, C. velutinum var. villosissimum
Name authority Linnaeus: Sp. Pl. 1: 440. (1753) Rafinesque: Med. Repos., hexade 2, 5: 359. (1808)
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