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common chickweed, common mouse-ear chickweed, céraiste commun, mouse-ear chickweed

alpine chickweed, Bering chickweed, Bering mouse-ear chickweed, Bering sea chickweed, Beringian chickweed, céraiste du détroit de Bering

Habit Plants perennial (rarely annual), tufted to mat-forming, often rhizomatous. Plants perennial, forming loose to dense mats or clumps, taprooted, with short, prostrate sterile shoots, rarely rhizomatous.
Stems

flowering stems erect from decumbent base, branched proximally, 10–45 cm, softly pubescent, eglandular with straight hairs; nonflowering shoots, when present, produced proximally, decumbent, rooting at nodes, branched, 5–20 cm, often subglabrous with alternating lines of eglandular hairs; small axillary tufts of leaves usually absent.

flowering stems usually erect, 10–25 cm, glabrous to sparsely pubescent proximally, more densely so in mid and distal stem, hairs patent to slightly deflexed, multicellular, glandular and eglandular;

internodes usually equaling or exceeding leaves; nonflowering shoots present; small axillary tufts of leaves absent.

Leaves

not marcescent;

blade 10–25(–40) × 3–8(–12) mm, densely covered with patent to ascending, colorless, long, eglandular hairs;

leaves of flowering shoots in distant pairs, sessile, blade elliptic to ovate-oblong, apex subacute;

leaves of sterile shoots pseudopetiolate, often spatulate, blade oblanceolate, apex obtuse.

sessile (flowering shoots) or petiolate (proximal stems and nonflowering shoots); flowering shoots with blade lanceolate, 5–20 × 2–5 mm, not succulent, apex ± acute, pubescent;

proximal stem and nonflowering shoots with blade oblanceolate often spatulate, not succulent, apex obtuse, ± pubescent on both surfaces (rarely subglabrous), with straight, strigose hairs or cilia, hairs pale, fuscous, mostly eglandular.

Inflorescences

lax, 3–50-flowered cymes;

bracts lanceolate, reduced, herbaceous, eglandular-pubescent, distal often with narrow, scarious margins.

open, dichotomous, (1–)3–10-flowered cymes;

bracts lanceolate, distal with scarious margins, proximal herbaceous and foliaceous, pubescence glandular and eglandular.

Pedicels

somewhat curved distally, 2–10(–20) mm, longer than sepals, densely pubescent with patent, eglandular, rarely glandular hairs.

erect, angled at base of calyx in fruit, 8–55 mm, 1–5 times as long as capsule, densely pubescent with patent glandular and eglandular hairs.

Flowers

sepals ovate-lanceolate, 5–7 mm, margins narrow, apex acute, scarious, pubescent with eglandular, rarely glandular, hairs;

petals oblanceolate, 1–1.5 times as long as sepals, apex deeply 2-fid;

stamens 10, occasionally 5;

styles 5.

sepals lanceolate to lance-elliptic, 3–7 mm, margins broad, apex ± acute, densely glandular-pubescent;

petals broadly oblanceolate, 6–12 mm, ± equaling (rarely to 2 times as long as) sepals, apex deeply 2-fid;

stamens 10;

styles 5.

Capsules

narrowly cylindric, curved, 9–17 mm, ca. 2 times sepals;

teeth 10, erect, margins convolute.

cylindric, curved, 8–12 mm, ca. 2 times sepals, rarely shorter;

teeth 10, erect, margins convolute.

Seeds

reddish brown, 0.4–1.2 mm, bluntly tuberculate;

testa not inflated, tightly enclosing seed.

pale to dark brown, 0.7–1.1 mm, tuberculate;

testa not inflated, tightly enclosing seed.

2n

= 122–152, usually 144.

= 72.

Cerastium fontanum

Cerastium beeringianum

Phenology Flowering spring and early summer.
Habitat Arctic and alpine tundra, meadows, open woodlands, rocky slopes, talus, cliff ledges, river and roadside gravel
Elevation 0-4000 m (0-13100 ft)
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK; YT; Europe [Introduced elsewhere]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (2 in the flora).

Cerastium fontanum is a highly variable and complex species. It often has been reported as C. vulgatum Linnaeus, an ambiguous name; see discussion under the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cerastium beeringianum is distinguished from C. alpinum by the absence of the long, silvery, flexuous, translucent, glistening hairs of that species. Cerastium beeringianum’s pubescence consists of straight, strigose, multicellular, somewhat fuscous hairs of several lengths, many of those in the mid and distal stem and inflorescence being glandular and viscid. The nodes and the leaves, at least in the mid and distal stem, typically have long, strigose, eglandular, fuscous hairs; those on the adaxial surface of the leaf being appressed, and those on the nodes retrorse. However, plants from the many small, isolated populations on the mountains of western North America show a great deal of variation. Some of these populations tend to be subglabrous, lacking most of the long hairs normally found on this species. Others are small, delicate plants with slender divaricate pedicels and smaller capsules and seeds. Though names have been given to several of these variants, they frequently intergrade, and much of the variation is greatly influenced by the environment.

Cerastium beeringianum is self-compatible and often self-pollinates, but the flowers are also freely visited by insects, particularly Diptera and smaller Hymenoptera, resulting in varying degrees of outbreeding.

Cerastium beeringianum intergrades with C. fischerianum, and it may be appropriate to treat them as subspecies of a single species. Unfortunately, the name C. fischerianum has priority.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences never glandular; petals 1.3-1.5 times as long as sepals; capsules 11-17 mm; seeds 0.9-1.2 mm
subsp. fontanum
1. Inflorescences occasionally viscid-glandular; petals equaling sepals; capsules 9-13 mm; seeds 0.4-0.9 mm
subsp. vulgare
Source FNA vol. 5. FNA vol. 5, p. 81.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Cerastium Caryophyllaceae > subfam. Alsinoideae > Cerastium
Sibling taxa
C. aleuticum, C. alpinum, C. arcticum, C. arvense, C. axillare, C. beeringianum, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. tomentosum, C. velutinum, C. viride
C. aleuticum, C. alpinum, C. arcticum, C. arvense, C. axillare, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. fontanum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. tomentosum, C. velutinum, C. viride
Subordinate taxa
C. fontanum subsp. fontanum, C. fontanum subsp. vulgare
Synonyms C. alpinum var. beeringianum, C. alpinum var. capillare, C. beeringianum var. capillare, C. beeringianum var. glabratum, C. beeringianum var. grandiflorum, C. buffumiae, C. earlei, C. fischerianum var. beeringianum, C. pilosum, C. pulchellum, C. scammamiae, C. variabile, C. vulgatum var. beeringianum
Name authority Baumgarten: Enum. Stirp. Transsilv. 1: 425. (1816) Chamisso & Schlechtendal: Linnaea 1: 62. (1826)
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