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céraiste des champs, field chickweed, field mouse-ear chickweed, field or prairie mouse-ear chickweed, meadow chickweed, starry cerastium

Photo is of parent taxon

céraiste dressé, field chickweed, field mouse-ear chickweed, Matted field chickweed

Habit Plants perennial, clumped and taprooted, or mat-forming and long-creeping rhizomatous. Plants forming clumps or mats, rhizomatous, or tufted, taprooted; straggling, creeping shoots usually not well developed.
Stems

flowering shoots often decumbent proximally, 5–20(–30) cm, glandular-pubescent distally, pilose-subglabrous, deflexed or spreading proximally;

non-flowering shoots present; small tufts of leaves present in axils of proximal leaves.

flowering stems decumbent at base, usually green or straw colored, occasionally purple tinged (in some populations growing on serpentine rocks), 5–20(–30) cm, pubescent and often glandular distally, hairs patent or deflexed; nonflowering winter shoots, when present, elongating, with narrow, oblanceolate leaves; small axillary tufts of leaves always present.

Leaves

not marcescent, sessile, ± spatulate proximally;

blade linear-lanceolate to lanceolate or narrowly oblong, 4–30 × 0.5–6 mm, apex acute, rarely obtuse, subglabrous to softly pubescent, sometimes glandular.

usually not strongly dimorphic;

blade lanceolate or oblanceolate to linear, 2–25 × 1–5 mm.

Inflorescences

lax, 1–20-flowered cymes, pubescence short, glandular;

bracts lanceolate, margins narrow, scarious, glandular-pubescent.

Pedicels

curved just below calyx, 5–30 mm, 1–6 times as long as sepals, glandular-pubescent.

Flowers

sepals narrowly lanceolate to lance-elliptic, 3.5–7 mm, margins narrow, softly pubescent;

petals obovate, 7.5–12.5 mm, ca. 2 times as long as sepals, apex 2-fid;

stamens 10;

anthers 0.8–1.1 mm;

styles 5.

sepals 3.5–6(–7) mm, midrib visible;

petals 7.5–9 mm (–12 mm in western plants), usually remaining ± white when dried;

anthers 0.8–0.9 mm.

Capsules

cylindric, curved, 7.5–11.5 × 2.5–4 mm, (1–)1.5–2 times as long as sepals;

teeth 10, erect, margins convolute.

7.5–11 × 2.5–4 mm, usually less than 1.5 times as long as sepals, rarely longer.

Seeds

brown, 0.6–1.2 mm diam., tuberculate;

testa not inflated.

0.6–1.1 mm.

2n

= 36, 72, (108, Europe).

= 36.

Cerastium arvense

Cerastium arvense subsp. strictum

Phenology Flowering spring.
Habitat Prairie grasslands, roadsides, arctic and alpine tundra, shores, dunes and rocky plains, rocky outcrops, alvars, sea cliffs and banks, favoring neutral to alkaline soils
Elevation 0-3800 m (0-12500 ft)
Distribution
from FNA
AK; AZ; CA; CO; CT; IA; ID; IN; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Worldwide
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; IA; ID; IN; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; SPM; Greenland; Europe (Alps); South America (s to Tierra del Fuego)
[BONAP county map]
Discussion

Subspecies 2+ (2 in the flora).

The infraspecific taxonomy of Cerastium arvense is subject to many different interpretations. While many subspecies have been recognized, the “actual” number is uncertain because of worldwide distribution, wide range of variation, and conflicting taxonomies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies strictum is widely distributed and grows in a great diversity of habitats, making it difficult to circumscribe and distinguish, both from subsp. arvense and from forms of Cerastium beeringianum, C. velutinum, and C. viride. Forms of subsp. strictum growing at high elevations or latitudes often develop broader leaves and may be confused with C. beeringianum (S. J. Wagstaff and R. J. Taylor 1988). However, C. arvense always has small axillary tufts of leaves. In northern parts of the Ungava region of Labrador, subsp. strictum appears on occasion to intergrade with C. alpinum. The status of these plants is uncertain.

Subspecies strictum is a remarkably variable taxon. Plants from the western side of the continent often have larger petals and a ranker growth. Completely glabrous plants (var. ophiticola) occur on serpentine in southern Quebec. Plants with broader ovate-elliptic leaves and tomentose pubescence (similar to Cerastium velutinum var. villosissimum but smaller) occur in the same area. Populations on the limestone plains near Belleville, southern Ontario, are more robust, with broader, strongly marcescent leaves at the base and a woolly pubescence. Plants from river valleys in Idaho tend to be much larger, with long, very narrow leaves; these are the basis for the name C. graminifolium. However, all of these plants are completely interfertile and show no reduction in fertility when crossed. Most of this variation is under genic control but also is affected by environmental factors.

Subspecies strictum is not interfertile with subsp. arvense or with other similar taxa such as Cerastium beeringianum, C. velutinum, and C. viride. Differences in chromosome numbers present an effective barrier to interfertility. However, several sterile hybrids involving subsp. strictum and those species have been synthesized. Many workers, most recently R. E. Ugborogho (1977), have included C. velutinum and C. viride in C. arvense as varieties or subspecies. However specific status is more appropriate because of the strong sterility barriers between them and the presence of morphological characters that enable them to be distinguished, albeit with difficulty in some herbarium material. This difficulty arises from the remarkable degree of variation displayed by subsp. strictum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Taproot absent, plant strongly rhizomatous with long-creeping shoots; flowering stems usually 25-30 cm, often purple pigmented proximally, pubescence eglandular (glandular hairs present in inflorescence), soft, short or subglabrous; sepals 5-7 mm; anthers 1-1.1 mm; petals usually turning brown when dried
subsp. arvense
1. Taprooted or shortly rhizomatous, forming clumps; flowering stems usually 5-20 cm, green or straw colored, glandular-pubescent; sepals 3.5-6(-7) mm; anthers 0.8-0.9 mm; petals usually remaining white when dried
subsp. strictum
Source FNA vol. 5, p. 79. FNA vol. 5, p. 80.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Cerastium Caryophyllaceae > subfam. Alsinoideae > Cerastium > Cerastium arvense
Sibling taxa
C. aleuticum, C. alpinum, C. arcticum, C. axillare, C. beeringianum, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. fontanum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. tomentosum, C. velutinum, C. viride
C. arvense subsp. arvense
Subordinate taxa
C. arvense subsp. arvense, C. arvense subsp. strictum
Synonyms C. alsophilum, C. angustatum, C. arvense var. angustifolium, C. arvense var. fuegianum, C. arvense var. latifolium, C. arvense var. ophiticola, C. arvense var. purpurascens, C. arvense var. sonnei, C. arvense var. strictum, C. arvense var. viscidulum, C. campestre, C. confertum, C. effusum, C. elongatum, C. fuegianum, C. graminifolium, C. leibergii, C. nitidum, C. occidentale, C. oreophilum, C. patulum, C. pensylvanicum, C. pubescens, C. scopulorum, C. sonnei, C. subulatum, C. tenuifolium, C. thermale, C. vestitum
Name authority Linnaeus: Sp. Pl. 1: 438. (1753) Gaudin: Fl. Helv. 3: 245. (1828)
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