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céraiste des champs, field chickweed, field mouse-ear chickweed, field or prairie mouse-ear chickweed, meadow chickweed, starry cerastium

Habit Plants perennial, clumped and taprooted, or mat-forming and long-creeping rhizomatous. Herbs, winter annual, annual, biennial, or perennial; taprooted and/or rhizomatous, rarely with tuberous thickenings (Pseudostellaria).
Stems

flowering shoots often decumbent proximally, 5–20(–30) cm, glandular-pubescent distally, pilose-subglabrous, deflexed or spreading proximally;

non-flowering shoots present; small tufts of leaves present in axils of proximal leaves.

prostrate to ascending or erect, simple or branched.

Leaves

not marcescent, sessile, ± spatulate proximally;

blade linear-lanceolate to lanceolate or narrowly oblong, 4–30 × 0.5–6 mm, apex acute, rarely obtuse, subglabrous to softly pubescent, sometimes glandular.

opposite, connate proximally or not, often petiolate (basal leaves), not stipulate;

blade subulate or linear to spatulate, lanceolate, or broadly ovate, seldom succulent.

Inflorescences

lax, 1–20-flowered cymes, pubescence short, glandular;

bracts lanceolate, margins narrow, scarious, glandular-pubescent.

terminal or axillary cymes, or flowers solitary;

bracts foliaceous or reduced, herbaceous to scarious (or rarely absent);

involucel bracteoles absent.

Pedicels

curved just below calyx, 5–30 mm, 1–6 times as long as sepals, glandular-pubescent.

present or rarely flowers sessile.

Flowers

sepals narrowly lanceolate to lance-elliptic, 3.5–7 mm, margins narrow, softly pubescent;

petals obovate, 7.5–12.5 mm, ca. 2 times as long as sepals, apex 2-fid;

stamens 10;

anthers 0.8–1.1 mm;

styles 5.

bisexual or seldom unisexual, sometimes inconspicuous;

perianth and androecium hypogynous or perigynous, often slightly;

hypanthium cup-, dish-, or disc-shaped;

sepals (4–)5, distinct or seldom connate basally, sometimes hooded, not awned;

petals absent or (1–)4–5, usually white, sometimes translucent, yellowish white, pink, or brownish, seldom clawed, auricles absent, coronal appendages absent, blade apex entire or 2-fid, sometimes jagged or emarginate, rarely laciniate;

stamens absent or (1–)5(–10), in 1 or 2 whorls, arising from base of ovary, a nectariferous disc, or sometimes the hypanthium or hypanthium rim;

staminodes absent or 1–5(–8);

ovary 1- or rarely 3-locular (Wilhelmsia);

styles (2–)3–5(–6), distinct;

stigmas (2–)3–5(–6).

Fruits

capsules, or rarely utricles (Scleranthus), opening by (2–)3–6, occasionally 8 or 10 valves or (3 or) 6–10 teeth;

carpophore present or often absent.

Capsules

cylindric, curved, 7.5–11.5 × 2.5–4 mm, (1–)1.5–2 times as long as sepals;

teeth 10, erect, margins convolute.

Seeds

brown, 0.6–1.2 mm diam., tuberculate;

testa not inflated.

1–60+, yellowish or tan to dark red or often brown or black, usually reniform or triangular to circular and laterally compressed or ovoid to globose, rarely oblong and dorsiventrally compressed (Holosteum);

embryo usually peripheral and curved, rarely central and straight (Holosteum).

x

= 6–15, 17–19, 23.

2n

= 36, 72, (108, Europe).

Cerastium arvense

Caryophyllaceae subfam. alsinoideae

Distribution
from FNA
AK; AZ; CA; CO; CT; IA; ID; IN; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; SD; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Worldwide
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North-temperate regions; South America (Andean region); Europe (Mediterranean region); w Asia; c Asia (Himalayas, Mediterranean region); Africa (Mediterranean region)
Discussion

Subspecies 2+ (2 in the flora).

The infraspecific taxonomy of Cerastium arvense is subject to many different interpretations. While many subspecies have been recognized, the “actual” number is uncertain because of worldwide distribution, wide range of variation, and conflicting taxonomies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 30, species ca. 1040 (16 genera, 137 species in the flora).

Alsinoideae, often considered basal in the family and the least specialized, is in some ways the most heterogeneous of the subfamilies. Members of its largest tribe (Alsineae) share the following characteristics: stipules absent, sepals free or at most basally connate, and capsular fruits. Indehiscent fruits, relatively short hypanthia, and other floral reductions occur in varying combinations in the approximately 30 species placed in four other tribes. A broad molecular survey of Alsinoideae has revealed two major lineages and lack of support for the existing tribal circumscriptions (M. Nepokroeff et al. 2002). About three-fourths of the species are members of Arenaria, Cerastium, Minuartia, and Stellaria.

Attempts have been made to move Scleranthus (fruit a utricle surrounded by an enlarged hypanthium) from Alsinoideae to either Paronychioideae (J. Hutchinson 1973, as Illecebraceae) or Scleranthaceae (A. Takhtajan 1997). Recent molecular and morphological studies by R. D. Smissen et. al. (2002, 2003) supported its retention in the Alsinoideae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Taproot absent, plant strongly rhizomatous with long-creeping shoots; flowering stems usually 25-30 cm, often purple pigmented proximally, pubescence eglandular (glandular hairs present in inflorescence), soft, short or subglabrous; sepals 5-7 mm; anthers 1-1.1 mm; petals usually turning brown when dried
subsp. arvense
1. Taprooted or shortly rhizomatous, forming clumps; flowering stems usually 5-20 cm, green or straw colored, glandular-pubescent; sepals 3.5-6(-7) mm; anthers 0.8-0.9 mm; petals usually remaining white when dried
subsp. strictum
Source FNA vol. 5, p. 79. FNA vol. 5, p. 50. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Cerastium Caryophyllaceae
Sibling taxa
C. aleuticum, C. alpinum, C. arcticum, C. axillare, C. beeringianum, C. bialynickii, C. brachypetalum, C. brachypodum, C. cerastoides, C. dichotomum, C. diffusum, C. dubium, C. fastigiatum, C. fischerianum, C. fontanum, C. glomeratum, C. maximum, C. nutans, C. pumilum, C. regelii, C. semidecandrum, C. terrae-novae, C. texanum, C. tomentosum, C. velutinum, C. viride
Subordinate taxa
C. arvense subsp. arvense, C. arvense subsp. strictum
Name authority Linnaeus: Sp. Pl. 1: 438. (1753) Fenzl: in S. L. Endlicher, Gen. Pl. 13: 963. (1840)
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