Cerastium alpinum
Caryophyllaceae subfam. alsinoideae
alpine chickweed, alpine mouse-ear chickweed, céraiste alpin
prostrate or ascending, tomentose (very rarely subglabrous), hairs white, translucent, long, soft, flexuous, some usually also short and glandular; flowering shoots ascending, 5–20 cm; small axillary tufts of leaves usually absent; nonflowering shoots ± prostrate, to 6 cm.
prostrate to ascending or erect, simple or branched.
marcescent or not, sessile;
blade obovate or ovate to elliptic-oblanceolate, elliptic, or lanceolate, usually 10–18 × 5–7 mm, apex obtuse, pubescence as on stems.
opposite, connate proximally or not, often petiolate (basal leaves), not stipulate;
blade subulate or linear to spatulate, lanceolate, or broadly ovate, seldom succulent.
open, (1–)2–4-flowered cymes;
bracts lanceolate, acute, margins narrow, scarious, glandular-pubescent.
terminal or axillary cymes, or flowers solitary;
bracts foliaceous or reduced, herbaceous to scarious (or rarely absent);
involucel bracteoles absent.
straight but often becoming angled at base and curved at apex, slender, 5–30 mm, often elongating to 3 or 4 times as long as sepals, pubescence usually dense, hairs both long, flexuous, multicellular, and short, glandular, viscid.
present or rarely flowers sessile.
sepals green, often violet tipped, narrowly elliptic-lanceolate, 7.5–10 mm, margins ± narrow, apex acute to obtuse, densely pubescent, hairs both long, eglandular and short, glandular;
petals 1–2 times as long as sepals, apex shallowly 2-fid;
stamens 10;
styles 5.
bisexual or seldom unisexual, sometimes inconspicuous;
perianth and androecium hypogynous or perigynous, often slightly;
hypanthium cup-, dish-, or disc-shaped;
sepals (4–)5, distinct or seldom connate basally, sometimes hooded, not awned;
petals absent or (1–)4–5, usually white, sometimes translucent, yellowish white, pink, or brownish, seldom clawed, auricles absent, coronal appendages absent, blade apex entire or 2-fid, sometimes jagged or emarginate, rarely laciniate;
stamens absent or (1–)5(–10), in 1 or 2 whorls, arising from base of ovary, a nectariferous disc, or sometimes the hypanthium or hypanthium rim;
staminodes absent or 1–5(–8);
ovary 1- or rarely 3-locular (Wilhelmsia);
styles (2–)3–5(–6), distinct;
stigmas (2–)3–5(–6).
capsules, or rarely utricles (Scleranthus), opening by (2–)3–6, occasionally 8 or 10 valves or (3 or) 6–10 teeth;
carpophore present or often absent.
cylindric, slightly curved, 12–16 mm, to 2 times as long as sepals;
teeth 10, erect, margins convolute.
dark brown, 1–1.4 mm diam., acutely tuberculate;
testa not inflated.
1–60+, yellowish or tan to dark red or often brown or black, usually reniform or triangular to circular and laterally compressed or ovoid to globose, rarely oblong and dorsiventrally compressed (Holosteum);
embryo usually peripheral and curved, rarely central and straight (Holosteum).
= 6–15, 17–19, 23.
= 72, 108.
Cerastium alpinum
Caryophyllaceae subfam. alsinoideae
Subspecies 3+ (2 in the flora).
The Cerastium alpinum group of species is a difficult complex of intergrading taxa. E. Hultén (1956) considered this complex to be the result of worldwide introgression among the various taxa. Members of this group in North America include C. aleuticum, C. alpinum, C. arcticum, C. beeringianum, C. bialynickii, C. fischerianum, C. regelii, and C. terrae-novae. Cerastium alpinum itself is distinguished from all other members of the complex by its lanate pubescence, which consists of long, silvery, translucent, multicellular, flexuous, often tangled hairs; the more or less square base of the calyx; the convex margins of the sepals; and, in well-grown plants, the long, slender, divaricate pedicels.
In western North America, Cerastium alpinum is replaced by C. beeringianum, which has long, straight, strigose, somewhat fuscous hairs, usually smaller flowers, and smaller seeds. The two species intergrade in eastern Canada; intermediate specimens were named C. alpinum var. strigosum Hultén.
Cerastium arcticum differs from C. alpinum, with which it often grows, in its straight, somewhat fuscous hairs; calyx which is round at the base; long, narrowly lanceolate sepals; large, straight, broad capsules; and broad, obtuse cauline leaves. Like C. alpinum, it usually has large flowers with the petals much longer than the sepals.
Many infraspecific taxa have been named in Cerastium alpinum but in North America it is much less variable than elsewhere. Two forms can be recognized at either the varietal or subspecific level.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 30, species ca. 1040 (16 genera, 137 species in the flora).
Alsinoideae, often considered basal in the family and the least specialized, is in some ways the most heterogeneous of the subfamilies. Members of its largest tribe (Alsineae) share the following characteristics: stipules absent, sepals free or at most basally connate, and capsular fruits. Indehiscent fruits, relatively short hypanthia, and other floral reductions occur in varying combinations in the approximately 30 species placed in four other tribes. A broad molecular survey of Alsinoideae has revealed two major lineages and lack of support for the existing tribal circumscriptions (M. Nepokroeff et al. 2002). About three-fourths of the species are members of Arenaria, Cerastium, Minuartia, and Stellaria.
Attempts have been made to move Scleranthus (fruit a utricle surrounded by an enlarged hypanthium) from Alsinoideae to either Paronychioideae (J. Hutchinson 1973, as Illecebraceae) or Scleranthaceae (A. Takhtajan 1997). Recent molecular and morphological studies by R. D. Smissen et. al. (2002, 2003) supported its retention in the Alsinoideae.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaf blades on flowering shoots narrowly elliptic or lance-elliptic to oblanceolate, apex ± acute, pubescence evenly distributed, not very dense; inflorescences usually (1-)2-4-flowered | subsp. alpinum |
1. Leaf blades on flowering shoots lance-elliptic to ovate or obovate, apex obtuse, pubescence a tuft of longer, silvery, ± tangled hairs at apex; inflorescences usually 1-flowered | subsp. lanatum |
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