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croix de malte, Maltese knapweed, Maltese star thistle or centaury, Maltese star-thistle, Napa thistle, tocalote

meadow knapweed, protean knapweed

Habit Annuals, 10–100 cm, herbage loosely gray-tomentose and villous with jointed multicellular hairs, sometimes minutely scabrous, minutely resin-gland-dotted. Perennials, 30–150 cm.
Stems

1–few, few–many branched distally.

1–few, erect or ascending, openly branched distally, villous to scabrous with septate hairs and loosely tomentose, ± glabrate.

Leaves

basal and proximal cauline petiolate or tapering to base, usually absent at anthesis, blades oblong to oblanceolate, 2–15 cm, margins entire to dentate or pinnately lobed;

cauline long-decurrent, blades linear to oblong or oblanceolate, 1–5 cm, entire or dentate.

basal and proximal cauline petiolate, blades oblanceolate or elliptic, 5–25 cm, margins entire or shallowly dentate to irregularly pinnately lobed;

distal cauline sessile, not decurrent, blades linear to lanceolate, gradually smaller, entire or dentate.

Involucres

ovoid, 10–15 mm, loosely cobwebby-tomentose or becoming glabrous.

ovoid to campanulate or hemispheric, 15–18 mm, usually ± as wide as high.

Florets

many;

corollas yellow, those of sterile florets 10–12 mm, slender, inconspicuous, those of fertile florets 10–12 mm.

40–100+, all fertile or the peripheral sterile;

corollas purple (rarely white), those of sterile florets ± expanded and exceeding corollas of fertile florets, those of fertile florets 15–18 mm.

Inner phyllaries

appendages entire, acute or spine-tipped.

tips truncate, irregularly dentate or lobed.

Heads

disciform, 1–few at branch tips, borne singly or in open leafy corymbiform arrays, sometimes clustered in distal axils, sessile or pedunculate.

usually radiant (rarely discoid), in few-headed corymbiform arrays, borne on leafy-bracted peduncles.

Cypselae

dull white or light brown, ca. 2.5 mm, finely hairy;

pappi of many white, unequal, stiff bristles 2.5–3 mm.

tan, 2.5–3 mm, finely hairy;

pappi 0 or of many unequal, sometimes caducous bristles 0.5–1 mm.

Principal

phyllaries: bodies ± stramineous, ovate, appendages purplish, spiny-fringed at base, each tipped by slender spine 5–10 mm.

phyllaries: bodies lanceolate to ovate, loosely tomentose or glabrous, usually concealed by expanded appendages, appendages erect, overlapping, light to dark brown, flat or ± concave, margins varying from coarsely dentate to pectinately dissected into ± wiry lobes.

2n

= 24.

= 22 (England), 44.

Centaurea melitensis

Centaurea ×moncktonii

Phenology Flowering mostly spring–summer (Apr–Jul). Flowering spring–fall (May–Nov).
Habitat Roadsides, fields, pine-oak woodlands, chaparral, agricultural areas Roadsides, riverbanks, pastures, meadows, forest openings, waste areas
Elevation 0–1500 m (0–4900 ft) 0–1000 m (0–3300 ft)
Distribution
from FNA
AL; AZ; CA; GA; ID; IL; MA; MO; MS; NJ; NM; NV; OR; PA; TX; UT; WA; WI; BC; Mexico (Baja California); Europe; Asia; Africa [Widely introduced]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CT; ID; IL; MA; ME; MI; MN; MO; MT; NH; NJ; NY; OH; OR; PA; RI; VA; VT; WA; WI; BC; NL; NS; ON; QC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Centaurea melitensis is native to the Mediterranean region. It is listed as a noxious weed in New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Centaurea ×moncktonii is native to Europe or originated in North America from European ancestry.

Meadow knapweeds represent an array of mutually interfertile intermediates derived by hybridization and backcrossing among the various cytotypes of the Centaurea jacea complex. The plants variously combine features of C. jacea and C. nigra, and perhaps C. nigrescens as well. The hybrid complex includes both diploids and tetraploids. Extremes approach the parental types. Meadow knapweeds are often present without either parent in the immediate vicinity. They are considered to be noxious weeds in British Columbia, Idaho, Oregon, and Washington.

Centaurea pratensis J. L. Thuillier, sometimes applied to plants that belong here, is not a legitimate name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 193. FNA vol. 19, p. 187.
Parent taxa Asteraceae > tribe Cardueae > Centaurea Asteraceae > tribe Cardueae > Centaurea
Sibling taxa
C. benedicta, C. calcitrapa, C. cyanus, C. depressa, C. diffusa, C. diluta, C. iberica, C. jacea, C. macrocephala, C. montana, C. nigra, C. nigrescens, C. phrygia, C. scabiosa, C. solstitialis, C. stoebe, C. sulphurea, C. virgata, C. ×moncktonii
C. benedicta, C. calcitrapa, C. cyanus, C. depressa, C. diffusa, C. diluta, C. iberica, C. jacea, C. macrocephala, C. melitensis, C. montana, C. nigra, C. nigrescens, C. phrygia, C. scabiosa, C. solstitialis, C. stoebe, C. sulphurea, C. virgata
Synonyms C. debeauxii subsp. thuillieri, C. jacea var. pratensis, C. jacea subsp. ×pratensis, C. nigra var. radiata, C. thuillieri
Name authority Linnaeus: Sp. Pl. 2: 917. (1753) C. E. Britton: Bot. Soc. Exch. Club Brit. Isles 6: 172. (1921)
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