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bois inconnu, common hackberry, hackberry, micocoulier occidental, western hackberry

bois inconnu, hackberry, sugarberry

Habit Trees or shrubs, size varying greatly in response to habitat; crowns rounded. Trees or rarely shrubs, to 30 m; crowns spreading.
Bark

gray, deeply furrowed, warty with age.

usually gray, smooth or often fissured and conspicuously warty.

Branches

without thorns, spreading, young branches mostly pubescent.

without or with thorns, slender, glabrous or pubescent.

Leaves

blade lance-ovate to broadly ovate or deltate, 5-12 × 3-6(-9) cm (on fertile branches), leathery, base oblique or obliquely somewhat acuminate, margins conspicuously serrate to well below middle, teeth 10-40, apex acuminate;

surfaces scabrous.

blade deltate to ovate to oblong-lanceolate, base oblique or cuneate to rounded, margins entire or serrate-dentate;

venation 3(-5)-pinnate.

Inflorescences

dense pendulous clusters.

staminate inflorescences cymes or fascicles; pistillate solitary or few-flowered clusters.

Flowers

usually unisexual, staminate and pistillate on same plants, along with a few bisexual flowers, pedicellate on branches of current year, appearing in mid or late spring.

Staminate flowers

filaments incurved in bud, exserted after anthesis;

gynoecium minute, rudimentary.

Pistillate flowers

calyx slightly to deeply 4(-5)-lobed;

stamens 4-5, inserted on pilose receptacle, included, often nonfunctional filaments usually shorter than in staminate flowers, rarely absent;

anthers ovate, face to face in bud, extrorse;

ovaries sessile, ovoid, 1-locular;

styles short, sessile, divided into 2 divergent, elongate, reflexed lobes, lobes entire or 2-cleft.

Fruits

fleshy drupes, ovoid or globose;

outer mesocarp thick, firm, inner mesocarp thin, fleshy;

stones thick walled, ripening in autumn, persisting after leaves fall.

Drupes

dark orange to purple- or blue-black when ripe, orbicular, to 7-11(-20) mm diam., commonly with thick beak;

pedicel to 15 mm.

Wood

light yellow, weak.

Stones

cream colored, 7-9 × 5-8 mm, reticulate.

x

= 10.

2n

= 20, 30, and 40.

Celtis occidentalis

Celtis

Phenology Flowering late winter–spring (Mar–May).
Habitat In rich moist soil along streams, on flood plains, on rock, on wooded hillsides, and in woodlands
Elevation 0-1800 m (0-5900 ft)
Distribution
from FNA
AL; AR; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; MB; ON; QC
[WildflowerSearch map]
[BONAP county map]
from USDA
Tropical and temperate regions; worldwide
[BONAP county map]
Discussion

Celtis occidentalis is valued as an ornamental street tree because of its tolerance to drought.

Native Americans used decoctions prepared from the bark of Celtis occidentalis medicinally as an aid in menses and to treat sore throat (D. E. Moerman 1986).

This is a highly variable species. Segregates named as varieties follow an east-west geographic gradient and are based primarily on leaf size, shape, and pubescence.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 60 (6 in the flora).

The hackberries provide important wildlife habitat, forming thickets that give shelter and fleshy drupes that ripen in autumn, persist after leaves fall, and supply winter food for birds and mammals. The treatment presented here is a simplified circumscription of species with no elaboration of infraspecific variation or interspecific hybridization. The group is taxonomically complex and in need of revision.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branches with thorns; leaf blade usually less than 2 cm wide.
C. pallida
1. Branches without thorns; leaf blade usually much more than 2 cm wide.
→ 2
2. Leaf blade typically elliptic-lanceolate to ovate-lanceolate, apex sharply acute to acuminate, margins mostly entire.
C. laevigata
2. Leaf blade typically broadly to narrowly ovate to oblong-lanceolate, apex blunt or obtuse to abruptly long-acuminate, acute, or short-acuminate, margins variable.
→ 3
3. Leaf blade abaxially white tomentose; fruits usually light brown; near San Antonio, Texas.
C. lindheimeri
3. Leaf blade abaxially essentially glabrous or with coarse pubescence mainly on veins; fruits mostly reddish orange to purple; widespread.
→ 4
4. Leaf blade typically 4.5 cm or less, margins entire or somewhat serrate above middle.
C. reticulata
4. Leaf blade mostly 5 cm or more, margins coarsely serrate for at least part of length.
→ 5
5. Shrubs or small trees; leaf blade serrate and sparingly toothed toward apex, entire proximally; fruits orange to brown to cherry red.
C. tenuifolia
5. Trees; leaf blade conspicuously serrate to well below middle; fruits dark orange to purple- or blue-black.
C. occidentalis
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ulmaceae > Celtis Ulmaceae
Sibling taxa
C. laevigata, C. lindheimeri, C. pallida, C. reticulata, C. tenuifolia
Subordinate taxa
C. laevigata, C. lindheimeri, C. occidentalis, C. pallida, C. reticulata, C. tenuifolia
Synonyms C. occidentalis var. canina, C. occidentalis var. crassifolia, C. occidentalis var. pumila, C. pumila, C. pumila var. deamii
Name authority Linnaeus: Sp. Pl. 2: 1044. (1753) Linnaeus: Sp. Pl. 2: 1043. 175: Gen. Pl. ed. 5, 467. (1754)
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