Flora of North America species comparison

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Ceanothus spinosus

Ceanothus papillosus

green-bark ceanothus, redheart

wartleaf ceanothus
Habit Shrubs, sometimes arborescent, evergreen, 2–6 m. Stems erect, not rooting at nodes; branchlets green, thorn-tipped or not, round to ± angled in cross section, rigid, glabrous or sparsely puberulent. Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose.
Leaves

not fascicled;

petiole 4–7 mm;

blade ± cupped, elliptic to oblong, 11–35 × 8–29 mm, base cuneate to rounded, margins entire, apex obtuse to weakly retuse, abaxial surface pale green, not glaucous, sometimes puberulent on midribs, adaxial surface green, shiny, glabrous; pinnately veined.

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.
Inflorescences

axillary, paniclelike, 4–17 cm.

axillary or terminal, racemelike, 2–8 cm.
Flowers

sepals, petals, and nectary pale blue to blue.

sepals, petals, and nectary deep blue.
Capsules

4–6 mm wide, not lobed;

valves smooth, not conspicuously viscid, not or weakly crested.

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.
2n

= 24.

= 24.

Ceanothus spinosus

Ceanothus papillosus
Phenology Flowering Jan–May. Flowering Mar–May.
Habitat Rocky slopes, coastal sage scrub, chaparral. Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests.
Elevation 60–900 m. (200–3000 ft.) 20–1500 m. (100–4900 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Like Ceanothus incanus, plants of C. spinosus are polymorphic for the presence of thorn-tipped branchlets. Putative hybrids with C. thyrsiflorus and C. oliganthus var. sorediatus have been reported (M. Van Rensselaer and H. McMinn 1944).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 12, p. 86. FNA vol. 12, p. 92.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 267. (1838) Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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