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wartleaf ceanothus

Sonoma ceanothus

Habit Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose. Shrubs, 0.5–1 m, often moundlike.
Stems

erect to ascending, not rooting at nodes;

branchlets gray to grayish brown, rigid, strigillose, glabrescent.

Leaves

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

not fascicled;

petiole 0–1 mm;

blade cupped, widely obovate to suborbiculate, 5–12 × 2–10 mm, base cuneate, margins not revolute, wavy, spinose-dentate, teeth 2–4, apex widely notched;

abaxial surface pale green or grayish green and glaucous, strigillose on veins, adaxial surface shiny green, glabrous.

Inflorescences

axillary or terminal, racemelike, 2–8 cm.

axillary or terminal, 0.8–1.5 cm.

Flowers

sepals, petals, and nectary deep blue.

sepals, petals, and nectary blue to lavender.

Capsules

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

4–5 mm wide, usually not, sometimes weakly lobed;

valves smooth, horns subapical, minute to ± prominent, erect, intermediate ridges absent.

2n

= 24.

= 24.

Ceanothus papillosus

Ceanothus sonomensis

Phenology Flowering Mar–May. Flowering Mar–Apr.
Habitat Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests. Sandy to rocky soils derived mostly from volcanic substrates, slopes, ridges, chaparral.
Elevation 20–1500 m. (100–4900 ft.) 100–700 m. (300–2300 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
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[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus sonomensis is distinctive in having spinose-dentate, few-toothed leaves, and slender fruit horns two to three millimeters; it occurs at a few scattered localities in the mountains of Napa and Sonoma counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 92. FNA vol. 12, p. 103.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Cerastes
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838) J. T. Howell: Leafl. W. Bot. 2: 162. (1939)
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