Flora of North America species comparison

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Ceanothus papillosus

Ceanothus incanus

wartleaf ceanothus

coast whitethorn
Habit Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose. Shrubs, evergreen, 1–4 m. Stems erect, not rooting at nodes; branchlets light gray, glaucous, thorn-tipped or not, round or slightly angled in cross section, rigid, puberulent, glabrescent.
Leaves

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

not fascicled;

petiole 3–12 mm;

blade flat, elliptic, ovate, or suborbiculate, 20–60 × 10–30 mm, base rounded or subcordate, margins entire, sometimes minutely glandular-denticulate above middle, glands 26–52, apex obtuse, abaxial surface pale green, appressed-puberulent, glabrescent, adaxial surface grayish green, dull, glabrate; 3-veined from base.
Inflorescences

axillary or terminal, racemelike, 2–8 cm.

axillary, usually paniclelike, sometimes racemelike, 3–6 cm.
Flowers

sepals, petals, and nectary deep blue.

sepals, petals, and nectary usually white to cream, sometimes pink.
Capsules

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

4–5 mm wide, ± lobed;

valves rugose, viscid when young, not or weakly crested.
2n

= 24.

= 24.

Ceanothus papillosus

Ceanothus incanus
Phenology Flowering Mar–May. Flowering Apr–Jun.
Habitat Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests. Flats, slopes, chaparral, open sites in conifer and mixed evergreen forests.
Elevation 20–1500 m. (100–4900 ft.) 60–1000 m. (200–3300 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus incanus is restricted to the Klamath and Santa Cruz mountains and North Coast Ranges of California. Some populations are evidently polymorphic for the presence of thorn-tipped or non-thorn-tipped branchlets (F. K. Klein 1970). Putative hybrids with C. papillosus and C. parryi have been reported (H. McMinn 1944); hybrids with C. thyrsiflorus have been called C. ×vanrensselaeri Roof.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 12, p. 92. FNA vol. 12, p. 86.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838) Torrey & A. Gray: Fl. N. Amer. 1: 265. (1838)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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