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wartleaf ceanothus

glory mat, Point Reyes ceanothus

Habit Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose. Shrubs, 0.1–3 m, matlike to moundlike.
Stems

prostrate, spreading, ascending, or erect, sometimes rooting at proximal nodes;

branchlets green to brown or reddish brown, flexible to rigid, strigillose or tomentulose.

Leaves

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

not fascicled, not crowded;

petiole 1–4 mm;

blade flat to ± cupped or folded lengthwise adaxially, widely elliptic, obovate, or suborbiculate, 10–40(–45) × 5–24 mm, base cuneate to ± rounded, margins not revolute, sometimes slightly thickened, dentate to denticulate most of length, teeth 9–35, apex rounded, truncate, or retuse, abaxial surface pale green, sparsely strigillose or glabrate, adaxial surface dark green, ± shiny, glabrous.

Inflorescences

axillary or terminal, racemelike, 2–8 cm.

axillary, 0.9–2.5 cm.

Flowers

sepals, petals, and nectary deep blue.

sepals, petals, and nectary deep blue to bluish purple.

Capsules

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

4–6 mm wide, lobed;

valves usually smooth, sometimes rugulose or crested distal to middle, horns subapical, minute, not rugose, intermediate ridges absent.

2n

= 24.

Ceanothus papillosus

Ceanothus gloriosus

Phenology Flowering Mar–May.
Habitat Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests.
Elevation 20–1500 m. (100–4900 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Ceanothus gloriosus is composed of three varieties occurring along the northern California coast from Humboldt County to Marin County. Variety gloriosus and var. porrectus generally differ primarily by leaf shape, length and width, and the number of marginal teeth. Variety exaltatus differs from the other two varieties primarily in stature. Complex hybrids with C. cuneatus var. ramulosus, C. divergens, and C. sonomensis were studied by J. T. Howell (1940).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Shrubs 0.8–3 m; stems erect to ascending.
var. exaltatus
1. Shrubs 0.1–0.5 m; stems prostrate to spreading.
→ 2
2. Leaf blades widely obovate to suborbiculate, 23–31(–45) × 17–24 mm, marginal teeth 13–31.
var. gloriosus
2. Leaf blades elliptic, obovate, or narrowly obovate, 10–21 × 5–15 mm, marginal teeth 9–19.
var. porrectus
Source FNA vol. 12, p. 92. FNA vol. 12, p. 102.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Cerastes
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Subordinate taxa
C. gloriosus var. exaltatus, C. gloriosus var. gloriosus, C. gloriosus var. porrectus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838) J. T. Howell: Leafl. W. Bot. 2: 43. (1937)
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