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wartleaf ceanothus

pine mat

Habit Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose. Shrubs, evergreen, 0.2–0.3(–0.5) m, matlike.
Stems

spreading, sometimes rooting at nodes;

branchlets green, sometimes tinged red, not thorn-tipped, usually round, sometimes angled, in cross section, flexible, puberulent.

Leaves

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

petiole 3–11 mm;

blade flat, elliptic to widely ovate, 12–45 × 6–20 mm, base obtuse to rounded, margins serrulate to denticulate, not revolute, usually not wavy, sometimes wavy, teeth 27–42, apex ± obtuse to rounded, abaxial surface pale green, pilosulous, adaxial surface green, pilosulous; usually pinnately veined, rarely 3-veined from base.

Inflorescences

axillary or terminal, racemelike, 2–8 cm.

axillary, umbel-like to racemelike, 1.3–4 cm.

Flowers

sepals, petals, and nectary deep blue.

sepals, petals, and nectary usually blue to pale blue, rarely white.

Capsules

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

4–5 mm wide, weakly lobed near apex;

valves smooth, crested.

2n

= 24.

Ceanothus papillosus

Ceanothus diversifolius

Phenology Flowering Mar–May. Flowering Apr–Jun.
Habitat Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests. Well-drained slopes and canyons, open to shaded sites, mixed evergreen and conifer forests.
Elevation 20–1500 m. (100–4900 ft.) 700–2300 m. (2300–7500 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus diversifolius occurs in the North Coast Ranges and the western slopes of the Cascade Range and the Sierra Nevada; it often forms mats to two meters wide. Marginal teeth on young leaves are notable in having more or less persistent, narrowly conic glands, not seen elsewhere in Ceanothus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 92. FNA vol. 12, p. 94.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus C. decumbens
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838) Kellogg: Proc. Calif. Acad. Sci. 1: 58. (1855)
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