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wartleaf ceanothus

mountain whitethorn, snow bush, whitethorn ceanothus

Habit Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose. Shrubs, evergreen, 0.5–1.5 m. Stems ascending to spreading, not rooting at nodes; branchlets yellowish or grayish green, glaucescent, thorn-tipped, round in cross section, rigid, puberulent, glabrescent.
Leaves

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

petiole 2–8 mm;

blade flat to cupped, ovate to elliptic, 10–30 × 6–18 mm, base rounded, margins usually entire, sometimes minutely glandular-denticulate distally, glands 18–30, apex obtuse, abaxial surface pale grayish green, sparsely puberulent or glabrous, sometimes villosulous along veins, adaxial surface pale green to grayish green, glaucous, dull, glabrate; 3-veined from base.

Inflorescences

axillary or terminal, racemelike, 2–8 cm.

axillary, umbel-like or racemelike, sometimes densely clustered, 1.2–2(–4) cm.

Flowers

sepals, petals, and nectary deep blue.

sepals, petals, and nectary usually white, rarely pink.

Capsules

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

3.5–5 mm wide, lobed;

valves rugose, viscid when young, weakly crested.

2n

= 24.

= 24.

Ceanothus papillosus

Ceanothus cordulatus

Phenology Flowering Mar–May. Flowering May–Jul.
Habitat Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests. Rocky ridges and slopes, chaparral, conifer and mixed evergreen forests.
Elevation 20–1500 m. (100–4900 ft.) 400–3400 m. (1300–11200 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus cordulatus is one of the most common shrubs in montane chaparral and forests of the Coast Ranges and Cascades of southern Oregon and northern California, southward through the Sierra Nevada, Transverse and Peninsular ranges of California, to the mountains of northern Baja California, and occurs disjunctly in the Charleston Mountains of Nevada.

Putative hybrids between Ceanothus cordulatus and C. velutinus var. velutinus, reported from the Klamath Mountains, the southern Cascade Range, and the Sierra Nevada, have been called C. ×lorenzenii (Jepson) McMinn. A rare intersectional hybrid between C. cordulatus and C. prostratus in the Lake Tahoe basin has been named C. ×serrulatus McMinn. Putative hybrids of C. cordulatus with C. diversifolius and C. integerrimus also have been reported (H. McMinn 1944).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 92. FNA vol. 12, p. 87.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838) Kellogg: Proc. Calif. Acad. Sci. 2: 124, fig. 39. (1863)
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