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deer brush

greasewood, mountain balm, snowbrush, snowbrush ceanothus, sticky-laurel, tobacco-brush, varnish-leaf ceanothus

Habit Shrubs, deciduous, 1.5–3 m. Stems ascending to erect, not rooting at nodes; branchlets pale green, not thorn-tipped, round in cross section, flexible, glabrous or strigillose. Shrubs, sometimes arborescent, evergreen, 1–6 m. Stems ascending to erect, not rooting at nodes; branchlets brown, not thorn-tipped, round in cross section, flexible or ± rigid, puberulent, glabrescent.
Leaves

not fascicled;

petiole 3–12 mm;

blade flat, lanceolate, elliptic, oblong-elliptic, or ovate, (10–)20–80 × 10–45 mm, base rounded, margins entire or denticulate distally, teeth 3–5, apex acute to obtuse, abaxial surface pale green, glabrate or puberulent, especially on veins, sometimes glabrescent, adaxial surface green, dull, glabrous or sparsely puberulent; usually 3-veined from base, rarely pinnately veined.

petiole 9–32 mm;

blade aromatic, flat, widely elliptic to ovate-elliptic, (25–)40–80(–130) × (13–)20–55(–60) mm, leathery, resinous, base subcordate to rounded, margins glandular-serrulate, teeth 93–150+, apex obtuse, abaxial surface pale green, velvety puberulent, especially on veins, or glabrous, adaxial surface dark green, shiny, glabrous; 3-veined from base.

Inflorescences

usually terminal, rarely axillary, racemelike or paniclelike, 5–25 cm.

axillary, paniclelike, 5–12 cm.

Flowers

sepals, petals, and nectary usually white or pale to dark blue, rarely pink.

sepals and petals cream;

nectary yellow-tinged.

Capsules

4–5 mm wide, lobed;

valves smooth, viscid, not or weakly crested.

3–4 mm wide, lobed at apex;

valves smooth or ± rugose, sometimes viscid, weakly crested or not crested.

2n

= 24.

Ceanothus integerrimus

Ceanothus velutinus

Phenology Flowering May–Jul.
Habitat Rocky slopes, open sites, chaparral, conifer and mixed evergreen forests.
Elevation 25–2600 m. (100–8500 ft.)
Distribution
from FNA
AZ; CA; NM; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; SD; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

G. T. Benson (1930) recognized four varieties in Ceanothus integerrimus, based on flower color, leaf blade shape, venation, and indumentum. With the exception of regional clines in indumentum density, N. C. Coile (1988) was unable to detect consistent patterns of variation correlated with ecological or geographical distribution. The types of C. integerrimus (D. Douglas in 1831) and C. andersonii (C. C. Parry in 1888) are notable in that they bear oblong leaf blades with one principal vein from the base; the basal pair of secondary veins are shorter than those distal to it and relatively short compared to those in leaves elsewhere in the species. This venation is restricted to only a few specimens from near Ben Lomond, Santa Cruz Mountains, California, and has not been seen since the collection by Parry. W. L. Jepson (1909–1943, vol. 2) suggested that the venation pattern may be correlated with the oblong leaf shape that is shown by the type specimens.

Young shoots of Ceanothus integerrimus were used by Native Americans in basketry and for arrow foreshafts (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

A common shrub on mountain slopes, Ceanothus velutinus, which reproduces by both seeds and layering, often forms large colonies, especially following fires or forest clearing. The leaves of C. velutinus are strongly aromatic (often vanilla-scented) when crushed, and the adaxial faces, especially in var. laevigatus, appear varnished.

An infusion of leaves of Ceanothus velutinus was used by Native Americans in cleansing and to treat skin inflammations (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade abaxial surfaces velvety puberulent, especially on veins; capsule valves smooth.
var. velutinus
1. Leaf blade abaxial surfaces glabrous, veins glabrous or sparsely puberulent; capsule valves ± rugose.
var. laevigatus
Source FNA vol. 12, p. 85. FNA vol. 12, p. 83.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. verrucosus
Subordinate taxa
C. velutinus var. laevigatus, C. velutinus var. velutinus
Synonyms C. andersonii, C. integerrimus var. californicus, C. integerrimus var. macrothyrsus, C. integerrimus var. puberulus
Name authority Hooker & Arnott: Bot. Beechey Voy., 329. (1838) Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 125, plate 45. (1831)
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