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deer brush

wartleaf ceanothus

Habit Shrubs, deciduous, 1.5–3 m. Stems ascending to erect, not rooting at nodes; branchlets pale green, not thorn-tipped, round in cross section, flexible, glabrous or strigillose. Shrubs, evergreen, 1–5 m. Stems erect to ascending, not rooting at nodes; branchlets green to reddish brown, not thorn-tipped, round in cross section, ± flexible to rigid, densely tomentulose.
Leaves

not fascicled;

petiole 3–12 mm;

blade flat, lanceolate, elliptic, oblong-elliptic, or ovate, (10–)20–80 × 10–45 mm, base rounded, margins entire or denticulate distally, teeth 3–5, apex acute to obtuse, abaxial surface pale green, glabrate or puberulent, especially on veins, sometimes glabrescent, adaxial surface green, dull, glabrous or sparsely puberulent; usually 3-veined from base, rarely pinnately veined.

petiole 1–3 mm;

blade cupped to flat, linear, narrowly oblong, or oblong-elliptic, 12–50 × 6–15 mm, base obtuse to rounded, margins minutely glandular-denticulate, revolute, glands 17–31, apex obtuse, truncate, or retuse, abaxial surface pale green, densely villosulous to tomentulose, adaxial surface dark green, sparsely puberulent and glandular-papillate; pinnately veined.

Inflorescences

usually terminal, rarely axillary, racemelike or paniclelike, 5–25 cm.

axillary or terminal, racemelike, 2–8 cm.

Flowers

sepals, petals, and nectary usually white or pale to dark blue, rarely pink.

sepals, petals, and nectary deep blue.

Capsules

4–5 mm wide, lobed;

valves smooth, viscid, not or weakly crested.

2–3 mm wide, lobed;

valves smooth, viscid when young, not or weakly crested.

2n

= 24.

= 24.

Ceanothus integerrimus

Ceanothus papillosus

Phenology Flowering May–Jul. Flowering Mar–May.
Habitat Rocky slopes, open sites, chaparral, conifer and mixed evergreen forests. Rocky ridges, slopes, and flats, chaparral, mixed evergreen forests.
Elevation 25–2600 m. (100–8500 ft.) 20–1500 m. (100–4900 ft.)
Distribution
from FNA
AZ; CA; NM; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
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[BONAP county map]
Discussion

G. T. Benson (1930) recognized four varieties in Ceanothus integerrimus, based on flower color, leaf blade shape, venation, and indumentum. With the exception of regional clines in indumentum density, N. C. Coile (1988) was unable to detect consistent patterns of variation correlated with ecological or geographical distribution. The types of C. integerrimus (D. Douglas in 1831) and C. andersonii (C. C. Parry in 1888) are notable in that they bear oblong leaf blades with one principal vein from the base; the basal pair of secondary veins are shorter than those distal to it and relatively short compared to those in leaves elsewhere in the species. This venation is restricted to only a few specimens from near Ben Lomond, Santa Cruz Mountains, California, and has not been seen since the collection by Parry. W. L. Jepson (1909–1943, vol. 2) suggested that the venation pattern may be correlated with the oblong leaf shape that is shown by the type specimens.

Young shoots of Ceanothus integerrimus were used by Native Americans in basketry and for arrow foreshafts (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ceanothus papillosus occurs in the Coast Ranges from San Francisco Bay south to the Santa Ynez Mountains, Ventura County, with disjunct populations in the Santa Ana Mountains, California, and Cerro Bola, in northern Baja California. The name C. papillosus var. roweanus was originally applied to low-growing plants with ascending to spreading, arcuate branches (H. McMinn 1939). M. Van Rensselaer and McMinn (1942) later emended the circumscription to include plants with linear leaves and retuse to truncate leaf apices, but these are found throughout the range of the species. Leaves with obtuse to truncate or retuse leaf apices also can be found on the same plant. Putative hybrids with C. integerrimus and C. oliganthus have been documented (McMinn 1944). Hybrids with C. thyrsiflorus have been named C. ×regius (Jepson) McMinn. Some putatively advanced generation hybrids have narrowly elliptic, weakly papillate leaves with obtuse apices, and sometimes have been misinterpreted as belonging to C. papillosus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 85. FNA vol. 12, p. 92.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Synonyms C. andersonii, C. integerrimus var. californicus, C. integerrimus var. macrothyrsus, C. integerrimus var. puberulus C. papillosus subsp. roweanus, C. papillosus var. roweanus
Name authority Hooker & Arnott: Bot. Beechey Voy., 329. (1838) Torrey & A. Gray: Fl. N. Amer. 1: 268. (1838)
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