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deer brush

hairy ceanothus, jimbrush (var. sorediatus)

Habit Shrubs, deciduous, 1.5–3 m. Stems ascending to erect, not rooting at nodes; branchlets pale green, not thorn-tipped, round in cross section, flexible, glabrous or strigillose. Shrubs, sometimes arborescent, evergreen, 2–3(–6) m. Stems erect, not rooting at nodes; branchlets reddish brown or brown, usually not, sometimes weakly, thorn-tipped, usually round, sometimes ± angled in cross section, flexible to rigid, glabrous, puberulent, or villosulous.
Leaves

not fascicled;

petiole 3–12 mm;

blade flat, lanceolate, elliptic, oblong-elliptic, or ovate, (10–)20–80 × 10–45 mm, base rounded, margins entire or denticulate distally, teeth 3–5, apex acute to obtuse, abaxial surface pale green, glabrate or puberulent, especially on veins, sometimes glabrescent, adaxial surface green, dull, glabrous or sparsely puberulent; usually 3-veined from base, rarely pinnately veined.

petiole 3–8 mm;

blade flat, ovate to widely elliptic, 8–35 × 4–25 mm, base obtuse to subcordate, margins denticulate most of length, not revolute, teeth 19–71, apex obtuse, rounded, or acute, abaxial surface usually pale green, sometimes gray-green, sometimes glaucous, glabrate to hirtellous, adaxial surface dark green, villosulous, especially on the veins, or glabrate; 3-veined from base.

Inflorescences

usually terminal, rarely axillary, racemelike or paniclelike, 5–25 cm.

axillary, racemelike, 1.5–5.2 cm.

Flowers

sepals, petals, and nectary usually white or pale to dark blue, rarely pink.

sepals, petals, and nectary usually pale to deep blue or purplish blue, rarely white.

Capsules

4–5 mm wide, lobed;

valves smooth, viscid, not or weakly crested.

4–7 mm wide, lobed, ± depressed at apex;

valves smooth or rugose, viscid, crested or not.

2n

= 24.

Ceanothus integerrimus

Ceanothus oliganthus

Phenology Flowering May–Jul.
Habitat Rocky slopes, open sites, chaparral, conifer and mixed evergreen forests.
Elevation 25–2600 m. (100–8500 ft.)
Distribution
from FNA
AZ; CA; NM; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

G. T. Benson (1930) recognized four varieties in Ceanothus integerrimus, based on flower color, leaf blade shape, venation, and indumentum. With the exception of regional clines in indumentum density, N. C. Coile (1988) was unable to detect consistent patterns of variation correlated with ecological or geographical distribution. The types of C. integerrimus (D. Douglas in 1831) and C. andersonii (C. C. Parry in 1888) are notable in that they bear oblong leaf blades with one principal vein from the base; the basal pair of secondary veins are shorter than those distal to it and relatively short compared to those in leaves elsewhere in the species. This venation is restricted to only a few specimens from near Ben Lomond, Santa Cruz Mountains, California, and has not been seen since the collection by Parry. W. L. Jepson (1909–1943, vol. 2) suggested that the venation pattern may be correlated with the oblong leaf shape that is shown by the type specimens.

Young shoots of Ceanothus integerrimus were used by Native Americans in basketry and for arrow foreshafts (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

M. Van Rensselaer and H. McMinn (1942) treated var. oliganthus and var. sorediatus as species, although they and R. F. Hoover (1970) discussed intergradation between the two forms throughout part of their distribution, especially in the southern Coast Ranges and Transverse Ranges of California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovaries and nectaries pilosulous; capsules glabrescent, valves rugose, weakly crested dorsally.
var. orcuttii
1. Ovaries and nectaries glabrous; capsules glabrous, valves smooth, crested or not.
→ 2
2. Branchlets puberulent to villosulous; leaf blade adaxial surfaces usually villosulous, sometimes glabrate; capsule valves usually crested, sometimes not.
var. oliganthus
2. Branchlets glabrous to sparsely puberulent; leaf blade adaxial surfaces sparsely villosulous to glabrate; capsule valves usually not, sometimes weakly, crested.
var. sorediatus
Source FNA vol. 12, p. 85. FNA vol. 12, p. 88.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Ceanothus
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Subordinate taxa
C. oliganthus var. oliganthus, C. oliganthus var. orcuttii, C. oliganthus var. sorediatus
Synonyms C. andersonii, C. integerrimus var. californicus, C. integerrimus var. macrothyrsus, C. integerrimus var. puberulus
Name authority Hooker & Arnott: Bot. Beechey Voy., 329. (1838) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 266. (1838)
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