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pine mat

mahala mat, prostrate ceanothus, squawcarpet

Habit Shrubs, evergreen, 0.2–0.3(–0.5) m, matlike. Shrubs, 0.1–0.3 m, matlike to moundlike.
Stems

spreading, sometimes rooting at nodes;

branchlets green, sometimes tinged red, not thorn-tipped, usually round, sometimes angled, in cross section, flexible, puberulent.

prostrate, spreading, or ascending, rooting at distal nodes;

branchlets reddish brown, ± flexible, puberulent, glabrescent.

Leaves

petiole 3–11 mm;

blade flat, elliptic to widely ovate, 12–45 × 6–20 mm, base obtuse to rounded, margins serrulate to denticulate, not revolute, usually not wavy, sometimes wavy, teeth 27–42, apex ± obtuse to rounded, abaxial surface pale green, pilosulous, adaxial surface green, pilosulous; usually pinnately veined, rarely 3-veined from base.

not fascicled, crowded, usually longer than internodes and obscuring them;

petiole 1–3 mm;

blade flat to ± cupped, elliptic to obovate, 6–30 × 4–16 mm, base cuneate, margins sometimes thick, not revolute, sometimes wavy, sharply dentate to spinose-dentate, teeth 3–9, apex rounded, abaxial surface pale green, glabrous except sparsely strigillose on veins, adaxial surface dark green, shiny, glabrate.

Inflorescences

axillary, umbel-like to racemelike, 1.3–4 cm.

axillary, 0.9–2 cm.

Flowers

sepals, petals, and nectary usually blue to pale blue, rarely white.

sepals, petals, and nectary pale to deep blue or purplish blue.

Capsules

4–5 mm wide, weakly lobed near apex;

valves smooth, crested.

6–9 mm wide, lobed;

valves smooth to rugulose, horns subapical, prominent, erect or spreading, rugose or not, intermediate ridges present.

2n

= 24.

Ceanothus diversifolius

Ceanothus prostratus

Phenology Flowering Apr–Jun.
Habitat Well-drained slopes and canyons, open to shaded sites, mixed evergreen and conifer forests.
Elevation 700–2300 m. (2300–7500 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Ceanothus diversifolius occurs in the North Coast Ranges and the western slopes of the Cascade Range and the Sierra Nevada; it often forms mats to two meters wide. Marginal teeth on young leaves are notable in having more or less persistent, narrowly conic glands, not seen elsewhere in Ceanothus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Putative hybrids between Ceanothus prostratus and C. velutinus var. velutinus, named C. ×rugosus, have been reported from northeastern California (H. McMinn 1944). A rare putative hybrid between C. prostratus and C. cordulatus in the Lake Tahoe basin has been named C. ×serrulatus. Both C. ×rugosus and C. ×serrulatus are intersubgeneric hybrids. Formally named hybrids between C. prostratus and C. cuneatus var. cuneatus include C. ×flexilis and possibly C. ×connivens, but the latter could have C. fresnensis as one of the parents rather than C. prostratus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades flat, margins not wavy, teeth usually 3–5(–7); capsule horns erect.
var. prostratus
1. Leaf blades slightly folded lengthwise adaxially, margins ± wavy, teeth 5–9; capsule horns spreading.
var. occidentalis
Source FNA vol. 12, p. 94. FNA vol. 12, p. 104.
Parent taxa Rhamnaceae > Ceanothus > subg. Ceanothus Rhamnaceae > Ceanothus > subg. Cerastes
Sibling taxa
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. prostratus, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
C. americanus, C. arboreus, C. arcuatus, C. confusus, C. cordulatus, C. crassifolius, C. cuneatus, C. cyaneus, C. dentatus, C. divergens, C. diversifolius, C. fendleri, C. ferrisiae, C. foliosus, C. fresnensis, C. gloriosus, C. griseus, C. hearstiorum, C. herbaceus, C. impressus, C. incanus, C. integerrimus, C. jepsonii, C. lemmonii, C. leucodermis, C. maritimus, C. martini, C. masonii, C. megacarpus, C. microphyllus, C. oliganthus, C. ophiochilus, C. otayensis, C. palmeri, C. papillosus, C. parryi, C. parvifolius, C. pauciflorus, C. perplexans, C. pinetorum, C. pumilus, C. purpureus, C. roderickii, C. sanguineus, C. sonomensis, C. spinosus, C. thyrsiflorus, C. tomentosus, C. velutinus, C. verrucosus
Subordinate taxa
C. prostratus var. occidentalis, C. prostratus var. prostratus
Synonyms C. decumbens
Name authority Kellogg: Proc. Calif. Acad. Sci. 1: 58. (1855) Bentham: Pl. Hartw., 302. (1849)
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