Caulanthus lemmonii |
Caulanthus |
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Lemmon's jewelflower, Lemmon's wild cabbage |
caulanthus, jewel-flower, wild-cabbage |
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Habit | Annuals; hispid basally, subglabrate or glabrous distally. | Annuals, biennials, or perennials; (caudex woody); not scapose; (usually glaucous), glabrous or pubescent, trichomes usually simple, rarely mixed with fewer, stalked, 2-rayed ones. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect or ascending, usually branched distally, 1–8 dm, sparsely hispid basally. |
usually erect or ascending, rarely subdecumbent, unbranched or branched distally, (sometimes inflated). |
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Leaves | basal and cauline; petiolate, subsessile, or sessile; basal rosulate or not, petiolate, blade margins entire, dentate, sinuate, lobed, or pinnatifid; cauline petiolate, subsessile, or sessile, blade (base sometimes sagittate or amplexicaul, rarely subauriculate), margins entire, dentate, or lobed. |
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Basal leaves | rosulate; petiole 0.3–3 cm; blade oblanceolate, 0.7–9 cm × 4–25 mm, margins coarsely dentate-sinuate. |
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Cauline leaves | (median) sessile; blade lanceolate to narrowly ovate, 0.5–11 cm × 2–45 mm, (smaller distally, base amplexicaul), margins entire or denticulate. |
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Racemes | (densely flowered), with a terminal cluster of sterile flowers. |
(corymbose, sometimes with a terminal cluster of sterile flowers from grouping of dark purple flower buds, rarely proximalmost flowers bracteate), elongated in fruit. |
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Flowers | sepals erect to ascending, (dark purple in bud, becoming greenish or creamy white and purplish or brown distally), ovate to narrowly lanceolate, 6–17 × 2.5–3.5 mm (subequal, keeled, usually glabrous, rarely pubescent, trichomes simple); petals white (with dark purple veins), 8–20 mm, blade 4–8 × 1.5–2 mm, crisped, claw oblanceolate, 4–11 × 2–3 mm; filaments in 3 unequal pairs, (median pairs often connate), abaxial pair 3.5–11mm, lateral pair 2–7 mm, adaxial pair 5–12 mm; anthers oblong to linear-oblong, unequal, 1.5–4 mm, (adaxial pair smaller). |
sepals usually erect or ascending, rarely spreading, oblong or ovate to lanceolate, (usually forming urceolate calyx, sometimes keeled), lateral pair saccate or not basally; petals yellow, brown, purple, lavender, white, yellowish green, or, rarely, pinkish, linear to oblanceolate, (often channeled and crisped, rarely neither, equal or abaxial (lower) pair slightly shorter than adaxial (upper) pair, longer than sepals), claw often well-differentiated from blade (sometimes longer and wider than blade, entire); stamens tetradynamous or 3 unequal pairs (rarely subequal); filaments not dilated basally, (usually distinct, sometimes adaxial or both median pairs connate); anthers oblong, (equal or adaxial pair smaller, apex obtuse or apiculate); nectar glands usually confluent, subtending bases of stamens. |
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Fruiting pedicels | ascending to divaricate, 3–18(–27) mm, pubescent or glabrous. |
erect, ascending to divaricate, or reflexed, slender or stout. |
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Fruits | erect or ascending (often straight), terete or slightly latiseptate, 5–12 cm × 2.5–3.5 mm; valves each with prominent midvein basally; ovules 52–72 per ovary; style 0–4 mm; stigma strongly 2-lobed (lobes 1–4 mm, opposite valves). |
sessile, linear, usually smooth, rarely torulose, usually terete, latiseptate, or angustiseptate, rarely 4-angled; valves each with distinct or obscure midvein, glabrous or sparsely pubescent; replum rounded; septum complete, (veinless); ovules 14–60[–210] per ovary; style obsolete or distinct; stigma capitate or not, entire, subentire, or 2-lobed (lobes opposite valves or replum). |
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Seeds | 2–3.5 × 1.7–2.2 mm. |
uniseriate, usually flattened (plump in C. californicus), usually not winged, rarely margined, oblong, ovoid, or, rarely, subglobose; seed coat not mucilaginous when wetted; cotyledons incumbent or obliquely so, (entire or, rarely, 3-fid). |
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x | = 14. |
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2n | = 28. |
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Caulanthus lemmonii |
Caulanthus |
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Phenology | Flowering (Feb-)Mar–May. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Grassland, chaparral, scrub | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 100-1100 m (300-3600 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA
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w United States; nw Mexico |
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Discussion | Of conservation concern. Both R. C. Rollins (1993) and R. E. Buck (1995) treated Caulanthus lemmonii as a variety of C. coulteri, whereas E. B. Payson (1923) treated the two as independent species. The differences between them clearly justify their separate recognition. In fact, those differences are far greater than those that distinguish the minor color form “barbarae” that both Rollins and Buck recognized as a distinct variety of C. amplexicaulis. The types of both C. coulteri and C. lemmonii are quite distinct. The slight intergradation between the two taxa, especially in occurrence of branched trichomes and lobing of cotyledons, most likely resulted from hybridization, but that needs to be verified experimentally and molecularly. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 17 (17 in the flora). The limits of Caulanthus have not changed for about 60 years following the excellent account by E. B. Payson (1923). R. C. Rollins (1993), however, resurrected Stanfordia mainly based on angustiseptate instead of terete fruits and on the presence of 3-fid cotyledons. The latter character is also present in C. coulteri, a species that Rollins maintained in Caulanthus. As emphasized by Payson and by I. A. Al-Shehbaz (1973), Stanfordia does not merit recognition, a position taken also by O. Appel and Al-Shehbaz (2003). R. E. Buck (1993, 1995) excluded three species of Caulanthus (C. anceps, C. flavescens, C. lasiophyllus) and placed them in the long-abandoned Guillenia. His primary reasoning was that they do not have the urceolate calyx or the crisped and channeled petals. However, C. flavescens has crisped and channeled petals, and sometimes has somewhat urceolate calyces, a feature not found in all species of Caulanthus. Extensive discussion on the maintenance of these three species in Caulanthus was given by Al-Shehbaz and needs not be repeated here. In my opinion, the recognition of Guillenia as distinct from Caulanthus is based on over-emphasis of a few petal characters and, without thorough molecular studies on the entire streptanthoid genera, it is more practical to maintain a more inclusive Caulanthus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 684. | FNA vol. 7, p. 677. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Caulanthus | Brassicaceae > tribe Thelypodieae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | C. coulteri var. lemmonii, Streptanthus coulteri var. lemmonii, Streptanthus parryi | Guillenia, Microsisymbrium, Stanfordia | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 23: 261. (1888) | S. Watson: Botany (Fortieth Parallel), 27, plate 3. (1871) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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