Caulanthus amplexicaulis |
Caulanthus crassicaulis |
|
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clasping leaf caulanthus, claspingleaf wild cabbage |
spindlestem, thick stem caulanthus, thick-stem wild cabbage |
|
Habit | Annuals; glabrous throughout. | Perennials; glabrous or sparsely pubescent. |
Stems | erect, unbranched or branched distally, 0.4–11 dm. |
erect or ascending, usually unbranched, rarely branched, (hollow, strongly inflated, fusiform, to 3 cm diam. at widest point), 2–10 dm. |
Basal leaves | rosulate; petiole 0.5–3 cm; blade obovate to broadly oblanceolate, 1–7 cm × 5–37 mm, margins coarsely dentate. |
rosulate; petiole 0.5–9 cm; blade obovate to oblanceolate (in outline), 1–12 cm × 3–45 mm, margins entire, dentate-sinuate, lyrate, runcinate, or pinnatifid. |
Cauline leaves | (median) sessile; blade oblong to broadly ovate or obovate, 2–8 cm × 10–40 mm (smaller distally, base amplexicaul), margins coarsely dentate or entire. |
(distalmost) shortly petiolate; blade linear to narrowly oblanceolate, margins entire. |
Racemes | (lax), without a terminal cluster of sterile flowers. |
(densely flowered), with a terminal cluster of sterile flowers. |
Flowers | sepals erect, (dark maroon-purple or ochroleucous to yellowish), ovate to ovate-lanceolate, 4–9.2 × 2–4 mm (equal); petals (abaxial pair) usually stramineous to yellowish green, rarely pale purple, (adaxial pair) purple, (slightly longer), 10–18 mm, blade 4–10 × 0.5–1.5mm, crisped, claw narrowly oblong-oblanceolate, 6–10 × 1–3 mm; filaments in 3 unequal pairs, abaxial pair 3–7 mm, lateral pair 1–5 mm, adaxial pair 4.5–9 mm; anthers oblong, unequal, 1–3.5 mm (adaxial pair smaller). |
sepals erect (creamy white, purplish, or greenish), ovate to lanceolate, 7.5–14 × 2.5–4 mm (equal); petals brown or purple, 10–15 mm, blade 3–5 × 1.2–2.5 mm, not or hardly crisped, claw oblanceolate, 7–10 × 2–2.5 mm; filaments tetradynamous, median pairs 3–8 mm, lateral pair 2–7 mm; anthers narrowly oblong, equal, 3–6.5 mm. |
Fruiting pedicels | divaricate to ascending, 2.5–19 mm. |
ascending, 1–5 mm, glabrous or pubescent, (trichomes flattened). |
Fruits | divaricate to ascending (often distinctly curved), terete or slightly latiseptate, 4.5–14(–16.7) cm × 1–1.5 mm; valves each with obscure midvein; ovules 40–92 per ovary; style 0–0.4 mm; stigma subentire. |
erect to ascending, terete or slightly latiseptate, 4.5–14 cm × 2–2.5 mm; valves each with obscure midvein; ovules 98–126 per ovary; style obsolete or, rarely, to 0.6 mm; stigma strongly 2-lobed (lobes to 1 mm, connivent, opposite valves). |
Seeds | 1.4–2.2 × 0.9–1.2 mm. |
1.5–4 × 1.5–2 mm. |
2n | = 28. |
= 28. |
Caulanthus amplexicaulis |
Caulanthus crassicaulis |
|
Phenology | Flowering Apr–Jul(-Aug). | Flowering Apr–Jul. |
Habitat | Chaparral, montane forests, serpentine areas, granitic and shale scree | Sagebrush scrub, pinyon-juniper woodland |
Elevation | 800-2900 m (2600-9500 ft) | 1200-2900 m (3900-9500 ft) |
Distribution |
CA
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AZ; CA; CO; ID; NV; UT; WY
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Discussion | Caulanthus amplexicaulis is known from Los Angeles, San Bernardino, Santa Barbara, and Ventura counties. The serpentine populations of Santa Barbara County have sepals that are ochroleucous to yellowish instead of the dark maroon ones found elsewhere in the species range. They were recognized by R. C. Rollins (1993) and R. E. Buck (1995) as var. barbarae. Both vars. barbarae and amplexicaulis form a distinct monophyletic clade but they are without reproductive barriers (A. E. Pepper and L. E. Norwood 2001). Some other species of Caulanthus show comparable or even greater variation in sepal color, and it might be more practical to treat var. barbarae as a color form. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
E. B. Payson (1923), R. C. Rollins (1993), and R. E. Buck (1995) recognized two varieties of Caulanthus crassicaulis and separated them solely on the basis of var. crassicaulis having pubescent sepals and glabrous or pubescent leaves, and var. glaber having glabrous or sparsely pubescent sepals and glabrous leaves. In my opinion, this distinction is impractical; as in some other species of Caulanthus, both glabrous and pubescent forms occur sometimes within the same population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 679. | FNA vol. 7, p. 681. |
Parent taxa | Brassicaceae > tribe Thelypodieae > Caulanthus | Brassicaceae > tribe Thelypodieae > Caulanthus |
Sibling taxa | ||
Synonyms | C. amplexicaulis var. barbarae, Euklisia amplexicaulis, Pleiocardia magna, Streptanthus amplexicaulis, Streptanthus amplexicaulis var. barbarae | Streptanthus crassicaulis, C. crassicaulis var. glaber, C. glaber, C. senilis |
Name authority | S. Watson: Proc. Amer. Acad. Arts 17: 364. (1882) | (Torrey) S. Watson: Botany (Fortieth Parallel), 27. (1871) |
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