Caulanthus |
Caulanthus cooperi |
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caulanthus, jewel-flower, wild-cabbage |
Cooper caulanthus, Cooper's jewel-flower, Cooper's wild cabbage |
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Habit | Annuals, biennials, or perennials; (caudex woody); not scapose; (usually glaucous), glabrous or pubescent, trichomes usually simple, rarely mixed with fewer, stalked, 2-rayed ones. | Annuals; puberulent or glabrous (trichomes simple and subappressed, and 2-rayed). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | usually erect or ascending, rarely subdecumbent, unbranched or branched distally, (sometimes inflated). |
erect to ascending (often flexuous, weak, often tangled with desert shrubs), usually branched distally, 1–8 dm, glabrous or puberulent. |
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Leaves | basal and cauline; petiolate, subsessile, or sessile; basal rosulate or not, petiolate, blade margins entire, dentate, sinuate, lobed, or pinnatifid; cauline petiolate, subsessile, or sessile, blade (base sometimes sagittate or amplexicaul, rarely subauriculate), margins entire, dentate, or lobed. |
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Basal leaves | rosulate; petiole 0.3–2.5 cm; blade oblanceolate to spatulate, 0.7–6 cm × 2–27 mm, margins usually coarsely dentate or somewhat pinnatifid, rarely entire, (surfaces glabrous). |
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Cauline leaves | (median) sessile; blade lanceolate or oblong, 1.5–7.5 cm × 5–20 mm (smaller distally, base amplexicaul to sagittate), margins dentate or entire, (surfaces glabrous). |
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Racemes | (corymbose, sometimes with a terminal cluster of sterile flowers from grouping of dark purple flower buds, rarely proximalmost flowers bracteate), elongated in fruit. |
(lax), without a terminal cluster of sterile flowers. |
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Flowers | sepals usually erect or ascending, rarely spreading, oblong or ovate to lanceolate, (usually forming urceolate calyx, sometimes keeled), lateral pair saccate or not basally; petals yellow, brown, purple, lavender, white, yellowish green, or, rarely, pinkish, linear to oblanceolate, (often channeled and crisped, rarely neither, equal or abaxial (lower) pair slightly shorter than adaxial (upper) pair, longer than sepals), claw often well-differentiated from blade (sometimes longer and wider than blade, entire); stamens tetradynamous or 3 unequal pairs (rarely subequal); filaments not dilated basally, (usually distinct, sometimes adaxial or both median pairs connate); anthers oblong, (equal or adaxial pair smaller, apex obtuse or apiculate); nectar glands usually confluent, subtending bases of stamens. |
sepals erect, (purplish or yellow-green), narrowly lanceolate, 3–6.5 × 0.8–1.5 mm (equal); petals yellow-green to purplish (often with purple veins), 4.5–9 mm, blade 2–3 × 0.7–1.5 mm, not crisped, claw narrowly oblong-oblanceolate, 2.5–7 × 1–1.5 mm; filaments slightly tetradynamous, median pairs 2–4.5 mm, lateral pair 1.5–3.5 mm; anthers oblong, equal, 1.5–2 mm. |
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Fruiting pedicels | erect, ascending to divaricate, or reflexed, slender or stout. |
reflexed, 1–4.5 mm, usually glabrous, rarely puberulent. |
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Fruits | sessile, linear, usually smooth, rarely torulose, usually terete, latiseptate, or angustiseptate, rarely 4-angled; valves each with distinct or obscure midvein, glabrous or sparsely pubescent; replum rounded; septum complete, (veinless); ovules 14–60[–210] per ovary; style obsolete or distinct; stigma capitate or not, entire, subentire, or 2-lobed (lobes opposite valves or replum). |
usually reflexed, rarely divaricate (often subfalcate), terete, 2–6 cm × 1.5–2.5 mm; valves each with prominent midvein, (glabrous or puberulent); ovules 24–48 per ovary; style 0.2–2.7 mm; stigma slightly 2-lobed. |
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Seeds | uniseriate, usually flattened (plump in C. californicus), usually not winged, rarely margined, oblong, ovoid, or, rarely, subglobose; seed coat not mucilaginous when wetted; cotyledons incumbent or obliquely so, (entire or, rarely, 3-fid). |
1–2 × 1–1.2 mm. |
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x | = 14. |
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2n | = 28. |
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Caulanthus |
Caulanthus cooperi |
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Phenology | Flowering (Jan-)Feb–Mar. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Desert shrubs, woodlands | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 600-2300 m (2000-7500 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
w United States; nw Mexico |
AZ; CA; NV; UT; Mexico (Baja California)
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Discussion | Species 17 (17 in the flora). The limits of Caulanthus have not changed for about 60 years following the excellent account by E. B. Payson (1923). R. C. Rollins (1993), however, resurrected Stanfordia mainly based on angustiseptate instead of terete fruits and on the presence of 3-fid cotyledons. The latter character is also present in C. coulteri, a species that Rollins maintained in Caulanthus. As emphasized by Payson and by I. A. Al-Shehbaz (1973), Stanfordia does not merit recognition, a position taken also by O. Appel and Al-Shehbaz (2003). R. E. Buck (1993, 1995) excluded three species of Caulanthus (C. anceps, C. flavescens, C. lasiophyllus) and placed them in the long-abandoned Guillenia. His primary reasoning was that they do not have the urceolate calyx or the crisped and channeled petals. However, C. flavescens has crisped and channeled petals, and sometimes has somewhat urceolate calyces, a feature not found in all species of Caulanthus. Extensive discussion on the maintenance of these three species in Caulanthus was given by Al-Shehbaz and needs not be repeated here. In my opinion, the recognition of Guillenia as distinct from Caulanthus is based on over-emphasis of a few petal characters and, without thorough molecular studies on the entire streptanthoid genera, it is more practical to maintain a more inclusive Caulanthus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Caulanthus cooperi is distributed in the Colorado and Mojave deserts in western Arizona, central and southern California, southern Nevada, and southern Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 677. | FNA vol. 7, p. 680. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Guillenia, Microsisymbrium, Stanfordia | Thelypodium cooperi, Guillenia cooperi | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | S. Watson: Botany (Fortieth Parallel), 27, plate 3. (1871) | (S. Watson) Payson: Ann. Missouri Bot. Gard. 9: 293. (1923) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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