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Sacramento Mountain Indian paintbrush, Wooton's paintbrush

Camp Martin paintbrush

Habit Herbs, perennial, 1.6–6.5 dm; from a woody caudex; with a woody taproot or branching roots. Herbs, perennial, 1.5–5(–8) dm; from a woody caudex; with a taproot.
Stems

solitary or few to many, erect, unbranched to much-branched, glabrous or hairy proximally and/or distally, hairs sparse to dense, spreading to erect, short to fairly long, soft, eglandular.

several to many, erect or ascending, sometimes curved at base, unbranched or branched, sometimes with short, leafy axillary shoots, hairs spreading, short or long and distally short, stiff or soft, stipitate-glandular.

Leaves

green, narrowly lanceolate to linear-lanceolate, linear, or narrowly elliptic, 2–8 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–5)-lobed, apex acuminate;

lobes widely spreading, linear to narrowly lanceolate, apex acute.

green to sometimes purple, narrowly oblong or lanceolate to sometimes linear or narrowly ovate, (0.8–)1–5(–6.5) cm, not fleshy, margins wavy, flat or involute, 0–5-lobed, apex rounded or obtuse, rarely acute;

lobes ascending-spreading or divaricate, linear to narrowly oblong, apex acute.

Inflorescences

3–16 × 2–3.5 cm;

bracts proximally greenish, distally red to orange-red, sometimes with a purplish medial band, lanceolate to broadly lanceolate or ovate in outline, (0–)3–5(–7)-lobed, sometimes also with 4 small teeth;

lobes ascending, linear-lanceolate, long, arising above or below mid length, apex acuminate, acute, or obtuse.

2.5–25 × 1.5–4 cm;

bracts proximally greenish, distally red, orange-red, or pale orange, rarely yellow, lanceolate or broadly lanceolate to oblong, (0–)3–5(–7)-lobed, plus 1–3 shallow teeth on apex of central lobe (var. clokeyi);

lobes ascending or spreading, linear, lanceolate, or oblanceolate, medium length or long, arising at or near mid length, central lobe apex obtuse to rounded, lateral ones acute.

Corollas

slightly to moderately curved distally, 25–37 mm;

tube 16–20 mm;

beak exserted 10–16 mm beyond calyx, adaxially green to yellowish, 11–25 mm;

abaxial lip green or red, small, inconspicuous, visible through cleft of calyx, 2 mm, 15–20% as long as beak;

teeth incurved, green or red, 0.7–1.5 mm.

straight to slightly curved, (19–)22–45 mm;

tube 11–22 mm;

abaxial lip rarely exserted, beak exserted;

beak adaxially green to yellowish, 9–21(–23) mm;

abaxial lip green or deep green with white teeth or red with green teeth, reduced, not inflated, often visible through front cleft, 1–2 mm, ca. 10% as long as beak;

teeth incurved, green to white, 0.5–1 mm.

Calyces

proximally green, distally red, 20–25 mm;

abaxial clefts 11–14(–17) mm, adaxial 8–9 mm, abaxial 50–60% of calyx length, adaxial 35–40% of calyx length, deeper than laterals, lateral 5–7 mm, 10–15% of calyx length;

lobes lanceolate to narrowly triangular, apex acute to acuminate.

proximally pale green, green, or whitish, sometimes with a yellow central band, distally colored as bracts, 14–26(–28) mm;

abaxial and adaxial clefts 3–10 mm, 20–50% of calyx length, deeper than laterals, lateral 1–5 mm, 15–20% of calyx length;

lobes lanceolate to broadly lanceolate, oblong, or ovate, apex acute, obtuse, or rounded.

2n

= 24, 48, 72.

Castilleja wootonii

Castilleja martini

Phenology Flowering Jun–Sep.
Habitat Grasslands, rocky slopes, ledges, canyons, open forests, montane to subalpine.
Elevation 2000–3700 m. (6600–12100 ft.)
Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; nw Mexico
Discussion

Castilleja wootonii is endemic to the White Mountains (Sierra Blanca) in south-central New Mexico and to the Mt. Livermore massif of western Texas. It should be sought in the intervening Guadalupe Mountains. Based on morphology, C. wootonii appears to be a southern derivative of C. linariifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Castilleja martini is sometimes treated as a subspecies of C. applegatei, but there are relatively consistent morphological discontinuities between these two variable complexes. See notes under 4. C. applegatei.

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes broadly lanceolate to ovate or oblong, apices acute, obtuse, or rounded, abaxial and adaxial clefts 20–44% of calyx lengths; w California.
var. martini
1. Calyx lobes lanceolate to broadly lanceolate, apices acute to sometimes obtuse, abaxial and adaxial clefts 40–50% of calyx lengths; sw Nevada, adjacent e California.
var. clokeyi
Source FNA vol. 17, p. 665. FNA vol. 17, p. 625.
Parent taxa Orobanchaceae > Castilleja Orobanchaceae > Castilleja
Sibling taxa
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. martini, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. xanthotricha
C. affinis, C. ambigua, C. angustifolia, C. applegatei, C. aquariensis, C. arachnoidea, C. attenuata, C. brevilobata, C. brevistyla, C. campestris, C. cervina, C. chambersii, C. chlorotica, C. christii, C. chromosa, C. chrymactis, C. chrysantha, C. cinerea, C. citrina, C. coccinea, C. collegiorum, C. covilleana, C. crista-galli, C. cryptantha, C. cusickii, C. densiflora, C. dissitiflora, C. disticha, C. elata, C. elegans, C. elmeri, C. exserta, C. flava, C. foliolosa, C. fraterna, C. genevieveana, C. glandulifera, C. gleasoni, C. gracillima, C. grisea, C. haydenii, C. hispida, C. hololeuca, C. hyperborea, C. indivisa, C. integra, C. kaibabensis, C. kerryana, C. kraliana, C. lacera, C. lanata, C. lasiorhyncha, C. lassenensis, C. latifolia, C. lemmonii, C. leschkeana, C. levisecta, C. linariifolia, C. lindheimeri, C. lineariloba, C. lineata, C. litoralis, C. lutescens, C. mendocinensis, C. mexicana, C. miniata, C. minor, C. mogollonica, C. mollis, C. montigena, C. nana, C. nelsonii, C. nervata, C. nivea, C. occidentalis, C. oresbia, C. organorum, C. ornata, C. pallescens, C. pallida, C. parviflora, C. parvula, C. patriotica, C. peckiana, C. peirsonii, C. pilosa, C. plagiotoma, C. praeterita, C. pruinosa, C. puberula, C. pulchella, C. purpurascens, C. purpurea, C. raupii, C. revealii, C. rhexiifolia, C. rigida, C. rubicundula, C. rubida, C. rupicola, C. salsuginosa, C. scabrida, C. schizotricha, C. septentrionalis, C. sessiliflora, C. subinclusa, C. suksdorfii, C. tenuiflora, C. tenuis, C. thompsonii, C. tomentosa, C. uliginosa, C. unalaschcensis, C. victoriae, C. viscidula, C. wightii, C. wootonii, C. xanthotricha
Subordinate taxa
C. martini var. clokeyi, C. martini var. martini
Synonyms C. ciliata C. applegatei subsp. martini
Name authority Standley: Muhlenbergia 5: 84. (1909) Abrams: Bull. S. Calif. Acad. Sci. 1: 69. (1902)
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