Castilleja viscidula |
Castilleja coccinea |
|
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sticky Indian paintbrush, sticky paintbrush |
painted-cup, red paintbrush, scarlet Indian paintbrush, scarlet paintbrush, scarlet painted-cup |
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Habit | Herbs, perennial, 0.5–3(–4) dm; from a woody caudex; with a taproot. | Herbs, annual or biennial, 1.2–5(–7) dm; with fibrous roots. |
Stems | several, ascending to erect, decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
solitary or few, erect to ascending, unbranched, rarely branched, hairs spreading, long, soft, eglandular, others shorter and glandular. |
Leaves | green to brown, linear, linear-lanceolate, lanceolate, or oblong (narrowly ovate nearing inflorescence), 1–4(–5) cm, not fleshy, margins wavy, flat or involute, (0–)2(–5)-lobed, apex acute; lobes ascending-spreading, oblong to narrowly lanceolate, apex acute or obtuse. |
green to sometimes brownish, rosette leaves persisting or withered at anthesis, oblanceolate to oblong-lanceolate, 1–3 cm, usually 0-lobed; cauline leaves linear-lanceolate to oblong-lanceolate, 2–7(–8) cm, not fleshy, margins plane, sometimes wavy, involute, 3–7(–9)-lobed, apex acute to acuminate; cauline leaves: lobes spreading to ascending, linear to oblong, (2–)5–25 mm, apex acuminate. |
Inflorescences | 2–14 × 1–3.5 cm; bracts proximally greenish to greenish brown, distally pale yellow, cream, or yellow-green, sometimes yellow-orange or red (sometimes gradually differentiated from proximal coloration), lanceolate, broadly lanceolate, or oblong, 3(–5)-lobed, proximal wavy-margined; lobes ascending, linear to narrowly lanceolate, long, arising near or above mid length, sometimes wavy-margined, apex acute to rounded. |
2–35 (longest in fruit) × 1.5–6 cm; bracts proximally greenish to brownish green, distally red, sometimes orange, yellow, or white, lanceolate to obovate, 3(–5)-lobed; lobes spreading-ascending, lanceolate and longer on proximal bracts, becoming oblanceolate, shorter, and wider on distal bracts, arising from distal 3/4 of blade, apex acute to rounded. |
Corollas | straight, 16–22(–25) mm; tube 10–15 mm; beak exserted, straight to sometimes curved, adaxially green to yellow, 5–8(–9) mm; abaxial lip green or yellow, sometimes deep purple, reduced, inconspicuous, often visible in abaxial cleft, 1–2 mm, 20% as long as beak; teeth erect, green to white, sometimes yellow or pink, 0.5–1 mm. |
straight, 18–31 mm; tube 15–18 mm; abaxial lip ± exserted, beak exserted, often whitish, yellowish, or faint dull reddish proximally, adaxially green distally, 7–10 mm; abaxial lip green to yellowish, apex pink to yellow, reduced, ± protuberant, 1.5–4 mm, 20–33% as long as beak; teeth erect, green or yellow, apices white, yellow, or pink, 2–4(–5) mm. |
Calyces | colored as bracts or proximally paler, (10–)14–18 mm; abaxial and adaxial clefts (4–)5–9 mm, 30–40% of calyx length, deeper than laterals, lateral (1–)2–6 mm, ca. 25% of calyx length; lobes narrowly ovate to lanceolate, linear, or narrowly lanceolate, apex acute to obtuse. |
colored as bracts, 17–23(–28) mm; abaxial clefts 6–10 mm, adaxial 5–8 mm, clefts 33–50% of calyx length, deeper than laterals, lateral 0–1 mm, 0–5% of calyx length; lobes oblong to triangular, apex rounded to truncate, sometimes emarginate. |
2n | = 24, 72. |
= 24, 46, 48. |
Castilleja viscidula |
Castilleja coccinea |
|
Phenology | Flowering Jun–Aug. | Flowering Jan–Sep. |
Habitat | Dry to mesic sagebrush steppes, rocky slopes, ledges, open woodlands, montane to subalpine. | Damp or wet meadows, roadsides, prairies, swamps, peatlands, ditches, thickets, dunes, jack pine flats, rocky forests, ledges, sandstone, limestone, or granite. |
Elevation | 2000–3200 m. (6600–10500 ft.) | 0–1200 m. (0–3900 ft.) |
Distribution |
ID; NV; OR
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AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; MB; ON; SK
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Discussion | Castilleja viscidula is a member of the complex including C. applegatei and C. martini, centered in California. Castilleja viscidula favors isolated mountain ranges, from the Wallowa and, possibly, the Blue mountains of northeastern Oregon, eastward into southwestern Idaho, and southward into central Nevada. Most populations are greenish yellow, but in one portion of the Wallowa Mountains, reddish bracted plants are common. Many yellowish bracted populations in the same mountain range surround this reddish population. Intermediate color forms are rarely encountered. Most ranges where C. viscidula occurs have generated slightly differing local races, demonstrating some reproductive isolation and divergence. In addition, hybrid swarms between this species and C. nana are known from several mountain ranges in central and northern Nevada, and an apparent hybrid with C. flava var. flava is known from the Independence Mountains of northern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja coccinea usually has red bracts. Forms with white or yellow bracts have been named but are scattered across the range of the species. However, yellow-bracted forms become markedly more common in populations in the northern portion of the range, especially in the upper midwestern region. Populations of C. coccinea seem ephemeral, disappearing from one site after a few years and appearing in another. It is rare in much of the eastern portion of its range, and apparently is extirpated in Louisiana, Maine, Massachusetts, and New Hampshire. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 664. | FNA vol. 17, p. 598. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | Bartsia coccinea, C. ludoviciana | |
Name authority | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 297. (1878) — (as Castilleia) | (Linnaeus) Sprengel: Syst. Veg. 2: 775. (1825) — (as Castilleia) |
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