Castilleja unalaschcensis |
Castilleja rubida |
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Alaska Indian paintbrush, coastal paintbrush, unalaska paintbrush |
little reddish Indian paintbrush, purple alpine paintbrush, purple paintbrush, Wallowa alpine paintbrush |
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Habit | Herbs, perennial, 2–8 dm; from a short, stout, scaly, many-headed, woody caudex; with rhizomes or branching, woody roots. | Herbs, perennial, 0.5–1.5 dm; from a woody caudex; with a taproot. |
Stems | few to many, erect to ascending, unbranched or branched distally, glabrate proximally, hairy distally, hairs sparse, spreading to erect, long, soft, eglandular, rarely stipitate-glandular. |
several, decumbent, or ascending, unbranched, hairs moderately dense, spreading, short and long, soft, eglandular and glandular. |
Leaves | green, sometimes red-purplish, lanceolate-linear to lanceolate, elliptic-ovate, or ovate, (1.2–)3.5–10 cm, not fleshy, margins plane, sometimes ± wavy, flat to slightly involute, 0(–3)-lobed, apex acuminate to acute; lobes ascending, lanceolate, apex acute to acuminate. |
green to purple, linear to narrowly lanceolate, 0.7–3.2 cm, not fleshy, margins plane, slightly involute, 3–5-lobed, apex narrowly acute to acuminate; lobes ascending-spreading, narrowly linear to filiform, often curling, often short, apex acute or obtuse. |
Inflorescences | 3.5–18 × 2–6.5 cm; bracts proximally green, distally yellow, pale yellow, whitish, yellow-green, or pale orange, often aging reddish or pink, lanceolate, elliptic, ovate, oblanceolate, or oblong to narrowly obovate, (0–)3–5-lobed; lobes ascending, linear to oblong, short, arising above mid length, apex acute to obtuse. |
2.5–6 × 1–2 cm; bracts purple, deep burgundy, or lavender throughout, rarely pink or yellowish white throughout, sometimes pink or dull whitish on distal margins and apices, oblong, 3–5(–7)-lobed; lobes spreading, linear, medium length, proximal lobes arising below mid length, center lobe apex rounded to obtuse, lateral ones acute to obtuse. |
Corollas | straight or slightly curved, 18–28 mm; tube 16–19 mm; beak partially to completely exserted, abaxial lip not exserted; beak adaxially green, 6–13 mm; abaxial lip deep green, reduced, inconspicuous, 1–2 mm, ca. 20% as long as beak; teeth incurved, deep green or white, 0.5–1 mm. |
straight, 12–15 mm; tube 14–16 mm; abaxial lip and beak exserted; beak adaxially green, 5–6 mm; abaxial lip colored as distal portion of bracts, prominent, pouches 3, central one grooved, pouches not strongly inflated, 4–5 mm, 80–100% as long as beak; teeth erect, appressed to beak, colored as distal portions of bracts, 1.5–2.5 mm. |
Calyces | colored as bracts, 15–28 mm; abaxial cleft 12–16 mm, adaxial 8–11.5 mm, clefts 40–50% of calyx length, deeper than laterals, lateral 4–10 mm, 20–35% of calyx length; lobes oblong to lanceolate, apex acute to obtuse or rounded. |
colored as bracts, 10–12 mm; all 4 clefts subequal, 3.5–6.5 mm, 35–55% of calyx length; lobes broadly linear or linear-triangular, apex obtuse to acute. |
Stigmas | green. |
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2n | = ca. 96. |
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Castilleja unalaschcensis |
Castilleja rubida |
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Phenology | Flowering May–Sep. | Flowering Jul–Aug. |
Habitat | Meadows, dunes, thickets, upper margins of tidal flats, river flats, gravel bars, tundra, open forests, roadsides, coastal to alpine. | Rocky slopes, ledges, dry to moist gravelly flats and ridges, alpine, limestone, rarely on river cobbles at lower elevations. |
Elevation | 0–2100 m. (0–6900 ft.) | 2200–3000 m. (7200–9800 ft.) |
Distribution |
AK; BC; YT; Asia (Russian Far East)
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OR |
Discussion | Castilleja unalaschcensis is a characteristic meadow species across much of the southern Alaskan coast, including the Aleutian Islands, as well as on the Queen Charlotte Islands (Haida Gwaii) and the mainland of western British Columbia and southern Yukon. It is common in the coastal littoral zone and ascends to lower alpine meadow communities. Morphologically, it appears to be a sister species to C. septentrionalis of the Rocky Mountains and boreal sites in eastern North America. A weakly differentiated interior form was described as subsp. transnivalis Pennell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja rubida is a rare alpine species endemic to a few limestone peaks in the Wallowa Mountains of northeastern Oregon, entirely within the Eagle Cap Wilderness Area. It is likely derived from the C. nana complex, found in the mountains of eastern California and Nevada, but it is amply distinct. Due to its very limited range and small population numbers, C. rubida is a species of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 663. | FNA vol. 17, p. 653. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. pallida var. unalaschcensis | |
Name authority | (Chamisso & Schlechtendal) Malte: Rhodora 36: 187. (1934) — (as unalaschkensis) | Piper: Bull. Torrey Bot. Club 27: 398. (1900) — (as Castilleia) |
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