Castilleja thompsonii |
Castilleja unalaschcensis |
|
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Thompson's Indian paintbrush, Thompson's owl clover, Thompson's paintbrush |
Alaska Indian paintbrush, coastal paintbrush, unalaska paintbrush |
|
Habit | Herbs, perennial, 0.8–4 dm; from a woody caudex; with a taproot. | Herbs, perennial, 2–8 dm; from a short, stout, scaly, many-headed, woody caudex; with rhizomes or branching, woody roots. |
Stems | few to many, erect or ascending, unbranched or branched, hairs spreading, long, stiff, sometimes soft (especially in higher elevations), eglandular, mixed with shorter stipitate-glandular ones. |
few to many, erect to ascending, unbranched or branched distally, glabrate proximally, hairy distally, hairs sparse, spreading to erect, long, soft, eglandular, rarely stipitate-glandular. |
Leaves | green to purple or reddish brown, linear to narrowly oblong or linear-lanceolate, 1.4–7.4 cm, not fleshy, margins plane to ± wavy, involute or flat, 3(–7)-lobed, apex acuminate; lobes spreading-ascending, linear, short to long moving up leaf axis, apex acute or obtuse. |
green, sometimes red-purplish, lanceolate-linear to lanceolate, elliptic-ovate, or ovate, (1.2–)3.5–10 cm, not fleshy, margins plane, sometimes ± wavy, flat to slightly involute, 0(–3)-lobed, apex acuminate to acute; lobes ascending, lanceolate, apex acute to acuminate. |
Inflorescences | 2.5–14 × 1–4 cm; bracts greenish to pale yellow or reddish brown throughout, or proximally greenish to dull reddish purple, or ruddy brown, distally greenish to yellow-green or yellow, often aging dull reddish to dull purplish, lanceolate to oblong to ovate, 3–5(–9)-lobed; lobes spreading to ascending, linear to narrowly lanceolate, long, proximal lobes arising below mid length, apex acute to obtuse. |
3.5–18 × 2–6.5 cm; bracts proximally green, distally yellow, pale yellow, whitish, yellow-green, or pale orange, often aging reddish or pink, lanceolate, elliptic, ovate, oblanceolate, or oblong to narrowly obovate, (0–)3–5-lobed; lobes ascending, linear to oblong, short, arising above mid length, apex acute to obtuse. |
Corollas | straight, 18–21 mm; tube 11–16 mm; subequal to calyx, sometimes beak exserted; beak adaxially green, 5–7(–8) mm; abaxial lip white, often proximally reddish, prominent, scarcely expanded, ± cylindric, 2.5–4(–5) mm, 50–70% as long as beak, glabrous or obscurely puberulent; teeth incurved to erect, white, 2.5 mm. |
straight or slightly curved, 18–28 mm; tube 16–19 mm; beak partially to completely exserted, abaxial lip not exserted; beak adaxially green, 6–13 mm; abaxial lip deep green, reduced, inconspicuous, 1–2 mm, ca. 20% as long as beak; teeth incurved, deep green or white, 0.5–1 mm. |
Calyces | colored as bracts, 12–25 mm; abaxial and adaxial clefts 4–8 mm, 20–60% of calyx length, deeper than laterals, lateral (0–)1–3 mm, 7–25% of calyx length; lobes linear, lanceolate, or triangular, apex acute, sometimes obtuse. |
colored as bracts, 15–28 mm; abaxial cleft 12–16 mm, adaxial 8–11.5 mm, clefts 40–50% of calyx length, deeper than laterals, lateral 4–10 mm, 20–35% of calyx length; lobes oblong to lanceolate, apex acute to obtuse or rounded. |
2n | = 24, 48. |
= ca. 96. |
Castilleja thompsonii |
Castilleja unalaschcensis |
|
Phenology | Flowering Apr–Aug(–Sep). | Flowering May–Sep. |
Habitat | Dry slopes, ridges, scabland lithosol soils, meadows, sagebrush steppes, valleys, montane to alpine. | Meadows, dunes, thickets, upper margins of tidal flats, river flats, gravel bars, tundra, open forests, roadsides, coastal to alpine. |
Elevation | 200–2100 m. (700–6900 ft.) | 0–2100 m. (0–6900 ft.) |
Distribution |
OR; WA; BC
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AK; BC; YT; Asia (Russian Far East)
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Discussion | Castilleja thompsonii is a characteristic species of the sagebrush communities on the eastern slope of the Cascade Range in Washington, and in the high deserts of the Columbia Basin. Historically, its range approached but apparently never entered Idaho in the Spokane River valley, but much of its habitat in that area is now converted to agriculture or suburban development or overwhelmed by non-native, invasive plants. Castilleja thompsonii occurs in a few sites in the Okanogan Valley region of southern British Columbia and at one site on the eastern slopes of the Cascade Range in Wasco County, Oregon. A distinctive form from the subalpine and alpine zones of Mt. Adams, in the southern Cascade Range of Washington, was named C. villicaulis. This form may merit varietal status under C. thompsonii. While both names were described in the same paper, C. thompsonii is the name used in all regional floras since their publication, after C. villicaulis was reduced to synonymy by M. Ownbey (1959). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja unalaschcensis is a characteristic meadow species across much of the southern Alaskan coast, including the Aleutian Islands, as well as on the Queen Charlotte Islands (Haida Gwaii) and the mainland of western British Columbia and southern Yukon. It is common in the coastal littoral zone and ascends to lower alpine meadow communities. Morphologically, it appears to be a sister species to C. septentrionalis of the Rocky Mountains and boreal sites in eastern North America. A weakly differentiated interior form was described as subsp. transnivalis Pennell. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 661. | FNA vol. 17, p. 663. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. villicaulis | C. pallida var. unalaschcensis |
Name authority | Pennell: Proc. Acad. Nat. Sci. Philadelphia 99: 178. (1947) — (as thompsoni) | (Chamisso & Schlechtendal) Malte: Rhodora 36: 187. (1934) — (as unalaschkensis) |
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