Castilleja thompsonii |
Castilleja fraterna |
|
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Thompson's Indian paintbrush, Thompson's owl clover, Thompson's paintbrush |
fraternal Indian paintbrush, fraternal paintbrush |
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Habit | Herbs, perennial, 0.8–4 dm; from a woody caudex; with a taproot. | Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with a taproot. |
Stems | few to many, erect or ascending, unbranched or branched, hairs spreading, long, stiff, sometimes soft (especially in higher elevations), eglandular, mixed with shorter stipitate-glandular ones. |
few to many, ± curved at base, ascending or erect distally, unbranched, hairs moderately dense, spreading, medium length and long, soft, mixed with shorter stipitate-glandular ones. |
Leaves | green to purple or reddish brown, linear to narrowly oblong or linear-lanceolate, 1.4–7.4 cm, not fleshy, margins plane to ± wavy, involute or flat, 3(–7)-lobed, apex acuminate; lobes spreading-ascending, linear, short to long moving up leaf axis, apex acute or obtuse. |
green, sometimes purple-tinged, lanceolate, broadly lanceolate, or narrowly ovate, 1–4.3 cm, not fleshy, margins plane, sometimes ± wavy, ± involute, 0–3-lobed, apex acute to acuminate; lobes ascending, often narrowly oblanceolate, apex acute to acuminate. |
Inflorescences | 2.5–14 × 1–4 cm; bracts greenish to pale yellow or reddish brown throughout, or proximally greenish to dull reddish purple, or ruddy brown, distally greenish to yellow-green or yellow, often aging dull reddish to dull purplish, lanceolate to oblong to ovate, 3–5(–9)-lobed; lobes spreading to ascending, linear to narrowly lanceolate, long, proximal lobes arising below mid length, apex acute to obtuse. |
3–8.5 × 1–4 cm; bracts greenish or bright to sometimes dull red, sometimes scarlet, orange, or pale pink throughout, or proximally greenish, distally as above, ovate to elliptic, (0–)3-lobed; lobes ascending, lanceolate to narrowly triangular, short, arising above mid length, apex acute to obtuse or acuminate. |
Corollas | straight, 18–21 mm; tube 11–16 mm; subequal to calyx, sometimes beak exserted; beak adaxially green, 5–7(–8) mm; abaxial lip white, often proximally reddish, prominent, scarcely expanded, ± cylindric, 2.5–4(–5) mm, 50–70% as long as beak, glabrous or obscurely puberulent; teeth incurved to erect, white, 2.5 mm. |
straight or slightly curved, 20–40 mm; tube 27 mm; abaxial lip often visible in abaxial cleft, sometimes exserted, beak usually exserted from calyx; beak adaxially green to yellow, 8–14 mm; abaxial lip red, black, green, white, or green and white, ± prominent, slightly rounded, 1.5–5 mm, ca. 33% as long as beak; teeth ascending, green, bright red, pink, white, or yellow, 0.5–3 mm. |
Calyces | colored as bracts, 12–25 mm; abaxial and adaxial clefts 4–8 mm, 20–60% of calyx length, deeper than laterals, lateral (0–)1–3 mm, 7–25% of calyx length; lobes linear, lanceolate, or triangular, apex acute, sometimes obtuse. |
brightly, conspicuously colored for at least distal 2/3, often throughout, providing much of inflorescence coloration, proximally pale whitish to pale pink, distally colored as in distal portion of bracts, sometimes colored throughout as in distal portion of bracts, 15–30 mm; abaxial and adaxial clefts 7–14 mm, 33–50% of calyx length, deeper than laterals, lateral 1–5 mm, 5–15% of calyx length; lobes narrowly to broadly triangular, apex rounded to acute. |
2n | = 24, 48. |
|
Castilleja thompsonii |
Castilleja fraterna |
|
Phenology | Flowering Apr–Aug(–Sep). | Flowering Jul–Aug(–Sep). |
Habitat | Dry slopes, ridges, scabland lithosol soils, meadows, sagebrush steppes, valleys, montane to alpine. | Moist or dry rocky slopes and flats, ridges, talus, dwarf willow mats, subalpine to alpine, rarely along stream channels at lower elevations, over sedimentary rocks, often limy. |
Elevation | 200–2100 m. (700–6900 ft.) | 2000–2900 m. (6600–9500 ft.) |
Distribution |
OR; WA; BC
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OR
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Discussion | Castilleja thompsonii is a characteristic species of the sagebrush communities on the eastern slope of the Cascade Range in Washington, and in the high deserts of the Columbia Basin. Historically, its range approached but apparently never entered Idaho in the Spokane River valley, but much of its habitat in that area is now converted to agriculture or suburban development or overwhelmed by non-native, invasive plants. Castilleja thompsonii occurs in a few sites in the Okanogan Valley region of southern British Columbia and at one site on the eastern slopes of the Cascade Range in Wasco County, Oregon. A distinctive form from the subalpine and alpine zones of Mt. Adams, in the southern Cascade Range of Washington, was named C. villicaulis. This form may merit varietal status under C. thompsonii. While both names were described in the same paper, C. thompsonii is the name used in all regional floras since their publication, after C. villicaulis was reduced to synonymy by M. Ownbey (1959). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja fraterna is endemic to one ridge system in the Wallowa Mountains of northeastern Oregon. A parallel ridge system in the range has a second Wallowa Mountains endemic, C. rubida. Castilleja fraterna is colored similarly to C. miniata, but the petaloid teeth of the abaxial corolla lip are more like C. chrysantha. It is possible that C. fraterna was derived through hybridization, though its chromosome number is unknown. It resembles the newly described C. kerryana in Montana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 661. | FNA vol. 17, p. 609. |
Parent taxa | Orobanchaceae > Castilleja | Orobanchaceae > Castilleja |
Sibling taxa | ||
Synonyms | C. villicaulis | |
Name authority | Pennell: Proc. Acad. Nat. Sci. Philadelphia 99: 178. (1947) — (as thompsoni) | Greenman: Bot. Gaz. 48: 147. (1909) |
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