Castilleja subinclusa |
Castilleja viscidula |
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Franciscan paint brush, longleaf Indian paintbrush |
sticky Indian paintbrush, sticky paintbrush |
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Habit | Herbs, perennial, 1.8–12 dm; from a woody caudex; with a taproot. | Herbs, perennial, 0.5–3(–4) dm; from a woody caudex; with a taproot. | ||||||||
Stems | solitary or few to many, erect to ascending, often leaning on nearby shrubs (var. jepsonii), unbranched or branched, often with small, leafy axillary shoots, hairs matted to spreading, short and long, soft, mixed with short stipitate-glandular ones. |
several, ascending to erect, decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
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Leaves | green, pale gray-green, sometimes dull reddish purple, linear to narrowly, sometimes broadly, lanceolate, 2–10.5 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–3)-lobed, apex acuminate to acute; lobes spreading, linear to short-lanceolate, often arising near mid length, apex acute to obtuse. |
green to brown, linear, linear-lanceolate, lanceolate, or oblong (narrowly ovate nearing inflorescence), 1–4(–5) cm, not fleshy, margins wavy, flat or involute, (0–)2(–5)-lobed, apex acute; lobes ascending-spreading, oblong to narrowly lanceolate, apex acute or obtuse. |
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Inflorescences | 3–30 × 1–7 cm; bracts green or dull purple throughout, or proximally green or dull purple, distally bright red to orange-red, sometimes salmon, orange, or yellow, usually linear to lanceolate, distal sometimes broadly lanceolate to oblong or elliptic, 0(–3)-lobed; lobes spreading to erect, linear to oblong or triangular, short or medium length, arising near or above mid length, apex acute to obtuse. |
2–14 × 1–3.5 cm; bracts proximally greenish to greenish brown, distally pale yellow, cream, or yellow-green, sometimes yellow-orange or red (sometimes gradually differentiated from proximal coloration), lanceolate, broadly lanceolate, or oblong, 3(–5)-lobed, proximal wavy-margined; lobes ascending, linear to narrowly lanceolate, long, arising near or above mid length, sometimes wavy-margined, apex acute to rounded. |
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Corollas | curved proximally, straight to slightly curved distally, 25–57 mm; tube 15–29 mm; beak subequal to or exceeding calyx but abaxial lip, beak, and often part of tube usually exserted out abaxial cleft; beak adaxially green, yellow-green, or yellow, 13–21 mm, margins red, orange, or yellow; abaxial lip green, yellow, reddish, blackish, or deep red-purple, reduced, protuberant, forming a platform, 1–2 mm, 5–25% as long as beak; teeth ascending, green, purple sometimes with reddish tip, yellow, or blackish, 0.5–1 mm. |
straight, 16–22(–25) mm; tube 10–15 mm; beak exserted, straight to sometimes curved, adaxially green to yellow, 5–8(–9) mm; abaxial lip green or yellow, sometimes deep purple, reduced, inconspicuous, often visible in abaxial cleft, 1–2 mm, 20% as long as beak; teeth erect, green to white, sometimes yellow or pink, 0.5–1 mm. |
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Calyces | proximally pale green, rarely pale purple, distally bright red or red-orange, rarely orange or yellow, or bright red or red-orange throughout, 20–42 mm; abaxial clefts 13–27 mm, adaxial 7–17 mm, abaxial 40–70% of calyx length, adaxial 12–33% of calyx length, deeper than laterals, lateral 0–7.5 mm, 0–35% of calyx length; lobes linear to narrowly triangular, strongly curved away from stem proximally and distally obviously curved toward stem, apex acute to acuminate to obtuse. |
colored as bracts or proximally paler, (10–)14–18 mm; abaxial and adaxial clefts (4–)5–9 mm, 30–40% of calyx length, deeper than laterals, lateral (1–)2–6 mm, ca. 25% of calyx length; lobes narrowly ovate to lanceolate, linear, or narrowly lanceolate, apex acute to obtuse. |
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2n | = 24, 48, 72, 96. |
= 24, 72. |
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Castilleja subinclusa |
Castilleja viscidula |
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Phenology | Flowering Jun–Aug. | |||||||||
Habitat | Dry to mesic sagebrush steppes, rocky slopes, ledges, open woodlands, montane to subalpine. | |||||||||
Elevation | 2000–3200 m. (6600–10500 ft.) | |||||||||
Distribution |
CA; nw Mexico
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ID; NV; OR
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Discussion | Varieties 3 (3 in the flora). Castilleja subinclusa is divided into three varieties with somewhat discontinuous ranges. Identification of the varieties is often difficult when comparing only a single trait but is more easily accomplished when the characters are considered in a suite. The three varieties are also separable by range and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja viscidula is a member of the complex including C. applegatei and C. martini, centered in California. Castilleja viscidula favors isolated mountain ranges, from the Wallowa and, possibly, the Blue mountains of northeastern Oregon, eastward into southwestern Idaho, and southward into central Nevada. Most populations are greenish yellow, but in one portion of the Wallowa Mountains, reddish bracted plants are common. Many yellowish bracted populations in the same mountain range surround this reddish population. Intermediate color forms are rarely encountered. Most ranges where C. viscidula occurs have generated slightly differing local races, demonstrating some reproductive isolation and divergence. In addition, hybrid swarms between this species and C. nana are known from several mountain ranges in central and northern Nevada, and an apparent hybrid with C. flava var. flava is known from the Independence Mountains of northern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 657. | FNA vol. 17, p. 664. | ||||||||
Parent taxa | ||||||||||
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Subordinate taxa | ||||||||||
Name authority | Greene: Pittonia 4: 2. (1899) — (as Castilleia) | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 297. (1878) — (as Castilleia) | ||||||||
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