Castilleja subinclusa |
Castilleja pruinosa |
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Franciscan paint brush, longleaf Indian paintbrush |
frosted Indian paintbrush, frosted paintbrush, pruinose paintbrush |
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Habit | Herbs, perennial, 1.8–12 dm; from a woody caudex; with a taproot. | Herbs to subshrubs, perennial, (1.2–)2–8 dm; from a woody caudex; with a taproot. | ||||||||
Stems | solitary or few to many, erect to ascending, often leaning on nearby shrubs (var. jepsonii), unbranched or branched, often with small, leafy axillary shoots, hairs matted to spreading, short and long, soft, mixed with short stipitate-glandular ones. |
several to many, erect or ascending, branched or unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs ± dense, spreading to appressed, whitish or ashy gray, long, soft, much-branched, rarely unbranched, eglandular or obscurely glandular. |
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Leaves | green, pale gray-green, sometimes dull reddish purple, linear to narrowly, sometimes broadly, lanceolate, 2–10.5 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–3)-lobed, apex acuminate to acute; lobes spreading, linear to short-lanceolate, often arising near mid length, apex acute to obtuse. |
green, linear-lanceolate to narrowly or broadly lanceolate, 2–8 cm, not fleshy, margins plane to sometimes wavy, flat or involute, 0-lobed, sometimes 3–5-lobed distally, apex obtuse to rounded, sometimes acute; lobes sometimes divergent, ascending-spreading, linear, apex acute. |
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Inflorescences | 3–30 × 1–7 cm; bracts green or dull purple throughout, or proximally green or dull purple, distally bright red to orange-red, sometimes salmon, orange, or yellow, usually linear to lanceolate, distal sometimes broadly lanceolate to oblong or elliptic, 0(–3)-lobed; lobes spreading to erect, linear to oblong or triangular, short or medium length, arising near or above mid length, apex acute to obtuse. |
3–20 × 1.5–4 cm; bracts proximally greenish, distally abruptly red, sometimes yellow, salmon, or orange-red, often with a narrow yellow medial band, lanceolate, broadly lanceolate, or oblong, 0–5-lobed; lobes spreading to ascending, linear, long, arising near mid length, apex obtuse to acute. |
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Corollas | curved proximally, straight to slightly curved distally, 25–57 mm; tube 15–29 mm; beak subequal to or exceeding calyx but abaxial lip, beak, and often part of tube usually exserted out abaxial cleft; beak adaxially green, yellow-green, or yellow, 13–21 mm, margins red, orange, or yellow; abaxial lip green, yellow, reddish, blackish, or deep red-purple, reduced, protuberant, forming a platform, 1–2 mm, 5–25% as long as beak; teeth ascending, green, purple sometimes with reddish tip, yellow, or blackish, 0.5–1 mm. |
slightly curved, 20–51 mm; tube 12–21 mm; beak long-exserted, adaxially green, yellow, or red, 12–33 mm; abaxial lip deep green, reduced, 1–2.5 mm, ca. 10% as long as beak; teeth incurved to erect, white, yellow, or green, 0.5–2 mm. |
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Calyces | proximally pale green, rarely pale purple, distally bright red or red-orange, rarely orange or yellow, or bright red or red-orange throughout, 20–42 mm; abaxial clefts 13–27 mm, adaxial 7–17 mm, abaxial 40–70% of calyx length, adaxial 12–33% of calyx length, deeper than laterals, lateral 0–7.5 mm, 0–35% of calyx length; lobes linear to narrowly triangular, strongly curved away from stem proximally and distally obviously curved toward stem, apex acute to acuminate to obtuse. |
proximally green to yellow, distally and abruptly colored as in distal bracts, often with a central yellow band, 13–28 mm; abaxial clefts 5–13 mm, adaxial 8–14 mm, clefts 33–70% of calyx length, deeper than laterals, lateral 3–9 mm, 20–25% of calyx length; lobes lanceolate to triangular, apex acute to acuminate. |
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2n | = 24, 48, 72, 96. |
= 48. |
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Castilleja subinclusa |
Castilleja pruinosa |
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Phenology | Flowering Mar–Aug. | |||||||||
Habitat | Dry flats, rocky slopes, talus, thickets, open forests and forest edges, usually over serpentine. | |||||||||
Elevation | 50–2600 m. (200–8500 ft.) | |||||||||
Distribution |
CA; nw Mexico
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CA; OR
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Discussion | Varieties 3 (3 in the flora). Castilleja subinclusa is divided into three varieties with somewhat discontinuous ranges. Identification of the varieties is often difficult when comparing only a single trait but is more easily accomplished when the characters are considered in a suite. The three varieties are also separable by range and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja pruinosa is a common and variable paintbrush at moderate elevations in the southern half of the Cascade Range, occurring westward in Oregon and in adjacent northwestern California. It is also found in the northern Sierra Nevada. The often dense pubescence of branched hairs distinguishes it from other similar species within its range. Castilleja gleasoni was sometimes included within C. pruinosa, but C. gleasoni is likely of hybrid origin and occurs only in the San Gabriel Mountains of southern California. There are significant morphological differences between C. gleasoni and C. pruinosa. Castilleja pruinosa forms occasional hybrids with C. applegatei var. pinetorum and with C. brevilobata. In some populations in Douglas and eastern Lane counties, Oregon, C. pruinosa forms introgressive swarms with C. hispida var. hispida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 657. | FNA vol. 17, p. 647. | ||||||||
Parent taxa | ||||||||||
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Name authority | Greene: Pittonia 4: 2. (1899) — (as Castilleia) | Fernald: Erythea 6: 50. (1898) — (as Castilleia) | ||||||||
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