Castilleja subinclusa |
Castilleja elata |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
Franciscan paint brush, longleaf Indian paintbrush |
Siskiyou Indian paintbrush, Siskiyou paintbrush, slender paintbrush |
|||||||||
Habit | Herbs, perennial, 1.8–12 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.9–6.2 dm; from a remote woody caudex; with a taproot. | ||||||||
Stems | solitary or few to many, erect to ascending, often leaning on nearby shrubs (var. jepsonii), unbranched or branched, often with small, leafy axillary shoots, hairs matted to spreading, short and long, soft, mixed with short stipitate-glandular ones. |
solitary, sometimes few, proximally creeping, becoming rhizomatous, distally ascending to erect, unbranched or branched, glabrous proximally, glabrate distally, hairs ascending, medium length, soft, eglandular. |
||||||||
Leaves | green, pale gray-green, sometimes dull reddish purple, linear to narrowly, sometimes broadly, lanceolate, 2–10.5 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–3)-lobed, apex acuminate to acute; lobes spreading, linear to short-lanceolate, often arising near mid length, apex acute to obtuse. |
widely spaced on stem, green, linear to lanceolate or narrowly oblong, 1.3–7.2 cm, not fleshy, margins plane, ± revolute, 0-lobed, apex broadly acute to acuminate, distalmost sometimes obtuse. |
||||||||
Inflorescences | 3–30 × 1–7 cm; bracts green or dull purple throughout, or proximally green or dull purple, distally bright red to orange-red, sometimes salmon, orange, or yellow, usually linear to lanceolate, distal sometimes broadly lanceolate to oblong or elliptic, 0(–3)-lobed; lobes spreading to erect, linear to oblong or triangular, short or medium length, arising near or above mid length, apex acute to obtuse. |
2–13.5 × 1.5–4.5 cm; bracts proximally greenish, rarely pink, distally pink, magenta, or purple, sometimes distally buff, dull yellow, cream, or light yellow-orange, ovate to elliptic or lanceolate, 0–5-lobed, sometimes also with 3 small teeth on center lobe; lobes ascending or ± spreading, narrowly lanceolate to oblong, medium length, arising near or above mid length, apex acuminate to acute, sometimes obtuse. |
||||||||
Corollas | curved proximally, straight to slightly curved distally, 25–57 mm; tube 15–29 mm; beak subequal to or exceeding calyx but abaxial lip, beak, and often part of tube usually exserted out abaxial cleft; beak adaxially green, yellow-green, or yellow, 13–21 mm, margins red, orange, or yellow; abaxial lip green, yellow, reddish, blackish, or deep red-purple, reduced, protuberant, forming a platform, 1–2 mm, 5–25% as long as beak; teeth ascending, green, purple sometimes with reddish tip, yellow, or blackish, 0.5–1 mm. |
straight, 15–25 mm; tube 7–12 mm; beak and abaxial lip exserted above calyx lobes or ± pendently exserted from abaxial calyx cleft; beak adaxially green to yellowish, 6–11 mm; abaxial lip white to green, reduced, inconspicuous, pouches 3, small, 0.5–1.5 mm, 10–15% as long as beak; teeth ascending, white to deep green, 0.5–1 mm. |
||||||||
Calyces | proximally pale green, rarely pale purple, distally bright red or red-orange, rarely orange or yellow, or bright red or red-orange throughout, 20–42 mm; abaxial clefts 13–27 mm, adaxial 7–17 mm, abaxial 40–70% of calyx length, adaxial 12–33% of calyx length, deeper than laterals, lateral 0–7.5 mm, 0–35% of calyx length; lobes linear to narrowly triangular, strongly curved away from stem proximally and distally obviously curved toward stem, apex acute to acuminate to obtuse. |
whitish, green, or pink, pale colored ones tending to age pink, lobes as in bracts, 9–17 mm; abaxial and adaxial clefts 4.5–8 mm, 50% of calyx length, deeper than laterals, lateral 2–3 mm, 17–22% of calyx length; lobes linear to oblanceolate, apex acute. |
||||||||
2n | = 24, 48, 72, 96. |
= 24. |
||||||||
Castilleja subinclusa |
Castilleja elata |
|||||||||
Phenology | Flowering May–Aug. | |||||||||
Habitat | Serpentine bogs and wetlands. | |||||||||
Elevation | 50–1900 m. (200–6200 ft.) | |||||||||
Distribution |
CA; nw Mexico
|
CA; OR
|
||||||||
Discussion | Varieties 3 (3 in the flora). Castilleja subinclusa is divided into three varieties with somewhat discontinuous ranges. Identification of the varieties is often difficult when comparing only a single trait but is more easily accomplished when the characters are considered in a suite. The three varieties are also separable by range and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja elata is endemic to the Siskiyou Mountains of northwestern California and adjacent Oregon. Although often treated as a subspecies of C. miniata, it differs from that species in its shorter corolla beaks and distinctive bimodal coloration, with some populations exclusively pale yellow to pale orange and others pink-purple to magenta, as well as its specialized habitat in serpentine wetlands, where it often grows alongside Darlingtonia californica. Castilleja miniata grows on more mesic to moderately xeric substrates in the general vicinity of C. elata but with no sign of intergradation between the two species. The origin and significance of the two discrete color forms of C. elata deserve further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||
Key |
|
|||||||||
Source | FNA vol. 17, p. 657. | FNA vol. 17, p. 604. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | C. miniata subsp. elata | |||||||||
Name authority | Greene: Pittonia 4: 2. (1899) — (as Castilleia) | Piper: Smithsonian Misc. Collect. 50: 201. (1907) | ||||||||
Web links |