Castilleja subinclusa |
Castilleja dissitiflora |
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Franciscan paint brush, longleaf Indian paintbrush |
Mount Hamilton paintbrush, Mt. Hamilton Indian paintbrush |
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Habit | Herbs, perennial, 1.8–12 dm; from a woody caudex; with a taproot. | Herbs, perennial, 1.8–4(–5) dm; from a woody caudex; with a taproot. | ||||||||
Stems | solitary or few to many, erect to ascending, often leaning on nearby shrubs (var. jepsonii), unbranched or branched, often with small, leafy axillary shoots, hairs matted to spreading, short and long, soft, mixed with short stipitate-glandular ones. |
several to many, erect to ascending, unbranched, sometimes branched, hairs spreading, long, soft, mixed with shorter stipitate-glandular ones. |
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Leaves | green, pale gray-green, sometimes dull reddish purple, linear to narrowly, sometimes broadly, lanceolate, 2–10.5 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0(–3)-lobed, apex acuminate to acute; lobes spreading, linear to short-lanceolate, often arising near mid length, apex acute to obtuse. |
green, linear to narrowly or broadly lanceolate, (1–)3–5(–6) cm, not fleshy, margins wavy (obscure on many pressed specimens), involute, usually 0–3(–5)-lobed, apex broadly acute to rounded; lobes widely spreading, linear to narrowly lanceolate, apex acute to acuminate. |
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Inflorescences | 3–30 × 1–7 cm; bracts green or dull purple throughout, or proximally green or dull purple, distally bright red to orange-red, sometimes salmon, orange, or yellow, usually linear to lanceolate, distal sometimes broadly lanceolate to oblong or elliptic, 0(–3)-lobed; lobes spreading to erect, linear to oblong or triangular, short or medium length, arising near or above mid length, apex acute to obtuse. |
2.5–10 × 2–5.5 cm; bracts proximally greenish, distally red to red-orange, narrowly lanceolate to broadly lanceolate, 3–5-lobed; lobes spreading, distal, if present, ascending, linear or narrowly oblanceolate to triangular, proximals long, arising below mid blade, distals short, sometimes mere teeth, near apex of central lobe, sometimes wavy-margined, apex obtuse to rounded, sometimes acute. |
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Corollas | curved proximally, straight to slightly curved distally, 25–57 mm; tube 15–29 mm; beak subequal to or exceeding calyx but abaxial lip, beak, and often part of tube usually exserted out abaxial cleft; beak adaxially green, yellow-green, or yellow, 13–21 mm, margins red, orange, or yellow; abaxial lip green, yellow, reddish, blackish, or deep red-purple, reduced, protuberant, forming a platform, 1–2 mm, 5–25% as long as beak; teeth ascending, green, purple sometimes with reddish tip, yellow, or blackish, 0.5–1 mm. |
straight to slightly curved, 24–38 mm; tube 14–21 mm; beak exserted from calyx, adaxially green, 11–16.5(–18) mm; abaxial lip green, reduced, visible or not through deep front cleft in calyx, 2 mm, 13% as long as beak; teeth incurved, green, 1 mm. |
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Calyces | proximally pale green, rarely pale purple, distally bright red or red-orange, rarely orange or yellow, or bright red or red-orange throughout, 20–42 mm; abaxial clefts 13–27 mm, adaxial 7–17 mm, abaxial 40–70% of calyx length, adaxial 12–33% of calyx length, deeper than laterals, lateral 0–7.5 mm, 0–35% of calyx length; lobes linear to narrowly triangular, strongly curved away from stem proximally and distally obviously curved toward stem, apex acute to acuminate to obtuse. |
whitish with green veins or green, sometimes purple, distally same color as bracts, sometimes with yellowish band below colored apices, 20–26(–29) mm; abaxial clefts (8–)13–16(–19) mm, adaxial 7–12(–14) mm, clefts 35–50% of calyx length, deeper than laterals, lateral 2–6(–8) mm, 10–30% of calyx length; lobes linear-lanceolate to lanceolate, apex obtuse to acute. |
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2n | = 24, 48, 72, 96. |
= 48. |
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Castilleja subinclusa |
Castilleja dissitiflora |
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Phenology | Flowering Jun–Aug. | |||||||||
Habitat | Sagebrush slopes often rocky, montane to subalpine. | |||||||||
Elevation | 1900–3300 m. (6200–10800 ft.) | |||||||||
Distribution |
CA; nw Mexico
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NV
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Discussion | Varieties 3 (3 in the flora). Castilleja subinclusa is divided into three varieties with somewhat discontinuous ranges. Identification of the varieties is often difficult when comparing only a single trait but is more easily accomplished when the characters are considered in a suite. The three varieties are also separable by range and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja dissitiflora is endemic to several mountain ranges in central and eastern Nevada, in the upper montane and lower subalpine zones. It has the deep abaxial calyx cleft of C. linariifolia and the stipitate-glandular, wavy-margined leaves of C. applegatei var. pinetorum. Based on morphological data, Holmgren suggested that it is an allopolyploid derived from hybridization of C. applegatei var. pinetorum and C. linariifolia. His proposal is plausible and should be further tested. Castilleja dissitiflora is a tetraploid, while both putative parental species have at least some diploid populations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 657. | FNA vol. 17, p. 603. | ||||||||
Parent taxa | ||||||||||
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Name authority | Greene: Pittonia 4: 2. (1899) — (as Castilleia) | N. H. Holmgren: Mem. New York Bot. Gard. 21(4): 46, figs. 6–8. (1971) | ||||||||
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